4. Dryopteris goldiana (Hook.) A. Gray (Goldie's fern) Pl.
6e,f; Map 24
Rhizome and petiole scales dark brown, concolorous or sometimes with lighter
margins, shiny, linear to ovate. Leaves 50–125 cm long, monomorphic, herbaceous
to papery. Leaf blades elliptic to somewhat obovate in outline, tapering
abruptly to the tip and broadest at or above the middle, pinnately compound
above to sometimes 2 times pinnately compound below, glabrous, flat. Pinnae
2–19 cm long, ovate-lanceolate to linear, the tips attenuate, the margins
shallowly toothed, the pinnules entire to deeply lobed. Basal lower segment of
basal pinna about as long as the basal upper segment and shorter than the
adjacent basal segment. Sori usually closer to the midribs than to the margins
of the pinnules or pinnule lobes. Indusia glabrous, thin, usually shriveling at
maturity. Spores 41–47 mm long. 2n=82. June–October.
Uncommon and widely scattered in Missouri,
mostly south of the Missouri River (northeastern U.S.
west to Iowa and south to Louisiana). Spring branches, mesic upland
forests of ravines and drainages, bottoms of sinkhole ponds, shaded sandstone
ledges, and shaded rock outcrops, on various substrates.
at the southwestern edge of this species’ distribution, which may explain the
highly disjunct, seemingly random pattern of localities and the broad range of
habitats in the state. For a discussion of problems in separating this species
from the closely related D. celsa, see the treatment of that species.
Also, for a brief mention of a potential hybrid with D. marginalis, see
the treatment of D. marginalis.