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Project Name Data (Last Modified On 6/5/2013)

Flora Data (Last Modified On 6/5/2013)
Species Mikania guaco H. & B.
PlaceOfPublication PI. Aeq. 2: 84. 1809.
Note TYPE: Colombia, Magdalena River between Mahates & Angostura, Humboldt & Bonpland (P, not seen, US, microfiche).
Synonym Mikania amara var. guaco (H. & B.) Baker in Mart., Fl. Bras. 6(2): 237. 1876. Willoughbya guaco (H. & B.) Kuntze, Rev. Gen. PI. 1: 372. 1891. Mikania olivacea Klatt, Bull. Soc. Roy. Bot. Belgique 31: 195. 1892 (1893). TYPE: Costa Rica, forets de Buenos-Aires, 2300 m, Pittier 4433 (BR?, not seen; GH?, not seen).
Description Coarse sparingly branched vines; stems mostly brownish, terete, striate, glabrate, usually fistulose. Leaves opposite; blades broadly ovate, to 25 cm long and 15 cm wide, the base short-acute to acuminate, the margins entire, the apex broadly to narrowly short-acuminate, the upper surface sparsely puberulous and with minute spicules, the lower surface rather densely short-puberulous with numerous yellowish glandular punctations, the veins mostly pinnate; petioles slender, to 4.5 cm long. Inflorescence rather densely corymbose with almost all heads in clusters of 3, the ultimate branches ca. 0-1 mm long. Heads ca. 8-10 mm high; subinvolucral bract linear-lanceolate, to 2 mm long, short-puberulous; involucral bracts oblong with rounded tips, 5-6 mm long, ca. 1 mm wide, the outer pair puberulous; corolla whitish, ca. 6 mm long, the tube narrow, 2.5-3.0 mm long, the limb narrowly funnelform, the lobes short-triangular, about as long as wide, with scattered slender hairs on the outer surface, the cells subquadrate and ca. 25 microns wide; anther appendages firm, triangular, about twice as long as wide; style base slender, usually sparsely papillose, the style appendages hirsute with many long papillae. Achenes with 5 ribs or grooves, thickly corticated and terete when fully mature, ca. 4 mm long, the sides sparsely puberulous; pappus of ca. 50 slender bristles which are not or only slightly enlarged apically.
Habit vines
Distribution widely distributed in tropical America from Mexico to Brazil
Note the most common representative of a group that has heads almost all in groups of 3-5, style branches hirsute with long papillae, and achenes highly corticated when fully mature. A close relative, M. parviflora (Aubl.) Karst., distinguished by its broad subinvolucral bracts, seems to occur as close to Panama as the Choco in Colombia. More distant geographically is RI. trinitaria DC. of Venezuela, Guayana, and adjacent Brazil. The latter has large bracts as in M. parviflora, but also has a deeply cleft corolla with long narrow lobes.
Specimen BOCAS DEL TORO: Vic. of Chiriqui Lagoon, Bustimentos "lighthouse," von Wedel 2920 (MO, US). CANAL ZONE: Chagres, Fendler 153 (MO, US). Vic. of Fort Sherman, Standley 30989 (US). CHIRIQUI: Trail from Paso Ancho to Monte Lirio, upper valley of Rio Chiriqui Viejo, 1500-2000 m, Allen 1578 (MO, US). 1 mi E of Cafias Gordas, near Costa Rican Border, 800-1200 m, Liesner 281 (MO). Pastures around El Boquete, 1000-1300 m, Pittier 2996 (US). DARIEN: Rio Sambu', 0.5 mi above Rio Venado, Duke 9292 (MO, OS, US). Asnati, 200-300 m, Duke 10050 (MO). Helipad, Camp Cafiasas, Upper Rio Sabana tributary, 312 ft, Duke 15547 (OS, REED). Along the Sambui River, Pittier 5544 (US). PANAMA: San Jose Island, Johnston 1372 (MO, US), 1279 (US), 1319 (MO, US). Cerro Campana, 2700 ft. McDaniel 6901 (FSU, MO).
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