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  • Reference article Brako, L. & J. L. Zarucchi. (eds.) 1993. Catalogue of the Flowering Plants and Gymnosperms of Peru. Monogr. Syst. Bot. Missouri Bot. Gard. 45: i–xl, 1–1286.
  • Reference article Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera. 2012. Rubiaceae a Verbenaceae. 4(2): i–xvi, 1–533. In G. Davidse, M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera (eds.) Fl. Mesoamer.. Missouri Botanical Garden Press, St. Louis.
  • Reference article Dodson, C.H. & A.H. Gentry. 1978. Flora of the Río Palenque Science Center: Los Ríos Province, Ecuador. Selbyana 4(1–6): i–xxx, 1–628.
  • Reference article Forzza, R. C. 2010. Lista de espécies Flora do Brasil http://floradobrasil.jbrj.gov.br/2010. Jardim Botânico do Rio de Janeiro, Rio de Janeiro.
  • Reference article Hokche, O., P. E. Berry & O. Huber. (eds.) 2008. Nuevo Cat. Fl. Vasc. Venez. 1–860. Fundación Instituto Botánico de Venezuela, Caracas.
  • Reference article Idárraga-Piedrahita, A., R. D. C. Ortiz, R. Callejas Posada & M. Merello. (eds.) 2011. Fl. Antioquia: Cat. 2: 9–939. Universidad de Antioquia, Medellín.
  • Reference article Jørgensen, P. M. & S. León-Yánez. (eds.) 1999. Cat. Vasc. Pl. Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: i–viii, 1–1181.
  • Reference article Killeen, T. J., E. García Estigarribia & S. G. Beck. (eds.) 1993. Guía Árb. Bolivia 1–958. Herbario Nacional de Bolivia & Missouri Botanical Garden, Edit. Quipus srl., La Paz.
  • Reference article Lorence, D. H. 1999. A nomenclator of Mexican and Central American Rubiaceae. Monogr. Syst. Bot. Missouri Bot. Gard. 73: 1–177. View in BotanicusView in Biodiversity Heritage Library
  • Reference article Renner, S. S., H. Balslev & L.B. Holm-Nielsen. 1990. Flowering plants of Amazonian Ecuador–A checklist. AAU Rep. 24: 1–241.
  • Reference article Terán Aguilar, J. 2012. Estruc. Compos. Parq. Nac. Carrasco 1–72. Tesis, Universidad Mayor de San Simón, Cochabamba.
 
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