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  • Reference article Forzza, R. C. 2010. Lista de espécies Flora do Brasil http://floradobrasil.jbrj.gov.br/2010. Jardim Botânico do Rio de Janeiro, Rio de Janeiro.
  • Reference article Funk, V. A., P. E. Berry, S. Alexander, T. H. Hollowell & C. L. Kelloff. 2007. Checklist of the plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contr. U.S. Natl. Herb. 55: 1–584. View in Biodiversity Heritage Library
  • Reference article Steyermark, J. A. 1972. Rubiaceae. 23: 227–832. In B. Maguire & Collaborators (eds.) Bot. Guyana Highl.—Part IX. Mem. New York Bot. Gard.. New York Botanical Garden Press, Bronx.
 
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