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  • Reference article Breedlove, D.E. 1986. Flora de Chiapas. Listados Floríst. México 4: i–v, 1–246.
  • Reference article Cowan, C. P. 1983. Flora de Tabasco. Listados Floríst. México 1: 1–123.
  • Reference article Foster, R. C. 1958. A catalogue of the ferns and flowering plants of Bolivia. Contr. Gray Herb. 184: 1–223. View in Biodiversity Heritage Library
  • Reference article Killeen, T. J. & T. S. Schulenberg. 1998. A biological assessment of Parque Nacional Noel Kempff Mercado, Bolivia. RAP Working Papers 10: 1–372.
  • Reference article Macbride, J. F. 1936. Rubiaceae, Flora of Peru. Publ. Field Mus. Nat. Hist., Bot. Ser. 13(6/1): 3–261.
  • Reference article Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118.
  • Reference article Moraes R., M. 1990. Lista preliminar de especies botánicas coleccionadas durante la Expedición Río Madre de Dios (Norte de Bolivia). Mus. Nac. Hist. Nat. (Bolivia) Com. 10: 32–52.
  • Reference article Nelson, C. H. 1978. Contribuciones a la flora de la Mosquitía, Honduras. Ceiba 22(1): 41–64.
  • Reference article Parker III, T. A. & B. Bailey. 1991. A biological assessment of the Alto Madidi region and adjacent areas of Northwest Bolivia. RAP Working Papers 1: 1–108.
  • Reference article Pérez J., L. A., M. Sousa Sánchez, A. M. Hanan-Alipi, F. Chiang Cabrera & P. Tenorio L. 2005. Vegetación terrestre. Cap. 4: 65–110. In J. Bueno, F Álvarez & S. Santiago (eds.) Biodivers. Tabasco. CONABIO-UNAM, México.
  • Reference article Standley, P. C. & L. O. Williams. 1975. Rubiaceae. 24(11): 1–274. In P. C. Standley & L. O. Williams (eds.) Fl. Guatemala, Part XI, Fieldiana, Bot.. Field Museum of Natural History, Chicago.
 
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