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  • Reference article Bacigalupo, N.M., E. L. Cabral & C.M. Taylor. 2008. Rubiaceae. 107(3): 2871–2920. In F. O. Zuloaga, O. N. Morrone & M. J. Belgrano (eds.) Cat. Pl. Vasc. Cono Sur, Monogr. Syst. Bot. Miss. Bot. Gard.. Missouri Botanical Garden, St. Louis.
  • Reference article Bernardi, L. 1985. Contribución a la dendrologia paraguaya: Secunda parte. Boissiera 37: 7–294.
  • Reference article Forzza, R. C. 2010. Lista de espécies Flora do Brasil http://floradobrasil.jbrj.gov.br/2010. Jardim Botânico do Rio de Janeiro, Rio de Janeiro.
  • Reference article Foster, R. C. 1958. A catalogue of the ferns and flowering plants of Bolivia. Contr. Gray Herb. 184: 1–223. View in Biodiversity Heritage Library
  • Reference article Funk, V. A., P. E. Berry, S. Alexander, T. H. Hollowell & C. L. Kelloff. 2007. Checklist of the Plants of the Guiana Shield (Venezuela: Amazonas, Bolivar, Delta Amacuro; Guyana, Surinam, French Guiana). Contr. U.S. Natl. Herb. 55: 1–584. View in Biodiversity Heritage Library
  • Reference article Hokche, O., P. E. Berry & O. Huber. (eds.) 2008. Nuevo Cat. Fl. Vasc. Venez. 1–860. Fundación Instituto Botánico de Venezuela, Caracas.
  • Reference article Killeen, T. J. & T. S. Schulenberg. 1998. A biological assessment of Parque Nacional Noel Kempff Mercado, Bolivia. RAP Working Papers 10: 1–372.
  • Reference article Steyermark, J. A. 1995. Flora of the Venezuelan Guayana Project.
  • Reference article Taylor, C.M. 2007. Palicourea Aubl. 5: 380–384. In M. G. L. Wanderly, G. J. Shepherd & A. M. Giulietti (eds.) Fl. Fanerog. Estado São Paulo. Editora Hucitec, São Paulo.
  • Reference article Taylor, C.M., J. A. Steyermark, P.G. Delprete, A. Vincentini, R. Cortés, D. C. Zappi, C. H. Persson, C. B. Costa & E. Anunciação. 2004. Rubiaceae. 8: 497–847. In J. A. Steyermark, P. E. Berry & B. K. Holst (eds.) Fl. Venez. Guayana. Missouri Botanical Garden Press, St. Louis.
  • Reference article Wood, J. R. I., F. Mamani M., P. Pozo, J. D. Soto & D. Villarroel Segarra. (eds.) 2011. Guía Darwin Pl. Chiquitania 1–212. Museo de Historia Natural Noel Kempff Mercado, Santa Cruz.
  • Reference article Zuloaga, F. O., O. N. Morrone, M. J. Belgrano, C. Marticorena & E. Marchesi. (eds.) 2008. Catálogo de las plantas vasculares del Cono Sur. Monogr. Syst. Bot. Missouri Bot. Gard. 107(1–3): i–xcvi, 1–3348.
 
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