| Colombia | | | Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera | 2012 | Verbenaceae | Fl. Mesoamer. | 4(2): 453–473 |
| Colombia | Antioquia | | Idárraga-Piedrahita, A., R. D. C. Ortiz, R. Callejas Posada & M. Merello | 2011 | Listado de las plantas vasculares del departamento de Antioquia | Fl. Antioquia: Cat. | 2: 9–939 |
| Costa Rica | | | Taylor, C.M., B. E. Hammel & W.C. Burger | 1991 | New species, combinations, and records in Rubiaceae from the La Selva Biological Station, Costa Rica | Selbyana | 12: 134–140 |
| Costa Rica | | | Lorence, D. H. | 1999 | A nomenclator of Mexican and Central American Rubiaceae | Monogr. Syst. Bot. Missouri Bot. Gard. | 73: 1–177 |
| Costa Rica | | | Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera | 2012 | Verbenaceae | Fl. Mesoamer. | 4(2): 453–473 |
| Costa Rica | Limón | | Lorence, D. H. | 1999 | A nomenclator of Mexican and Central American Rubiaceae | Monogr. Syst. Bot. Missouri Bot. Gard. | 73: 1–177 |
| Ecuador | | | Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera | 2012 | Verbenaceae | Fl. Mesoamer. | 4(2): 453–473 |
| Ecuador | Pichincha | | Jørgensen, P. M. & S. León-Yánez | 1999 | (none) | Cat. Vasc. Pl. Ecuador, Monogr. Syst. Bot. Miss. Bot. Gard. | 75: i–viii, 1–1181 |
| Nicaragua | | | Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera | 2012 | Verbenaceae | Fl. Mesoamer. | 4(2): 453–473 |
| Nicaragua | | | Lorence, D. H. | 1999 | A nomenclator of Mexican and Central American Rubiaceae | Monogr. Syst. Bot. Missouri Bot. Gard. | 73: 1–177 |
| Panama | | | Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera | 2012 | Verbenaceae | Fl. Mesoamer. | 4(2): 453–473 |
| Panama | Bocas del Toro | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Canal Area | | Lorence, D. H. | 1999 | A nomenclator of Mexican and Central American Rubiaceae | Monogr. Syst. Bot. Missouri Bot. Gard. | 73: 1–177 |
| Panama | Canal Area | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Chiriquí | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Colón | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Darién | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Panamá | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |
| Panama | Veraguas | | Correa A., M.D., C. Galdames & M. Stapf | 2004 | (none) | Cat. Pl. Vasc. Panamá | 1–599 |