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Accepted By:
  • Reference article Delprete, P.G. & R. Cortés-Ballén. 2016. Rubiaceae. 2: 2252–2343. In R. Bernal González, S. R. Gradstein & M. Celis (eds.) Cat. Pl. Lich. Colombia. Universidad Nacional de Colombia (Sede Bogotá), Instituto de Ciencias Naturales, Bogotá, Colombia.
  • Reference article Jørgensen, P. M. & S. León-Yánez. (eds.) 1999. Cat. Vasc. Pl. Ecuador. Monogr. Syst. Bot. Missouri Bot. Gard. 75: i–viii, 1–1181.
 
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