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Published In: Species Muscorum Frondosorum 47. 1801. (Sp. Musc. Frond.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 9/27/2009)
Acceptance : Accepted
Project Data     (Last Modified On 10/8/2017)
Plant Category: Mosses
General Reference:

Notes     (Last Modified On 10/8/2017)

A genus typical of high elevations in the Neotropics, mostly associated with páramo and puna in open sites on exposed rocks. Andreaea is stated to contain nearly 100 species of cold temperate regions; Murray (1988, see family reference) has suggested that 50 species is a more realistic estimate. Although more than 25 species are listed for the tropical Andes, it is estimated that there are 10-15. The genus requires a critical revision for the Neotropics where numerous species have been described. Species provisionally excluded for which the report or taxonomic status is unknown is given below.


Un género típico de grandes elevaciones en el Neotrópico, generalmente asociado con rocas expuestas en sitios abiertos del páramo y la puna. Se afirma que Andreaea contiene casi 100 especies en regiones templadas frías; Murray (1988, ver referencias de la familia) sugirió que 50 especies es una estimación más realista. Sin embargo, más de 25 especies son listadas para los Andes tropicales, se estima que hay 15-20 especies. El género requiere una revisión crítica para el Neotrópico donde numerosas especies han sido descritas.


Status Unknow:


Andreaea angustifolia Broth., Biblioth. Bot. 87: 9. 1916. Type: Bolivia. Huaillattani, Quimzacruz, alt.4000 m, Herzog 2965.


Andreaea barbuloides Broth., Biblioth. Bot. 87: 10. 1916. Type: Bolivia. Hochtal Viloco, Quimzacruz, alt. 4600 m, Herzog 3158.


Andreaea clavata Broth., Biblioth. Bot. 87: 10. 1 f. 4. 1916. Type: Bolivia. Felsen am Huaillattanisee, alt. 4900 m, Herzog 2958. Paratype cited: Tunari, alt. 5000 m, Herzog 4785.


Andreaea clavata var. tenuior Broth., Bot. 87: 10. 1916. Type: Bolivia. Huaillattani, Quimzacruz, Herzog 2960.


Andreaea dissitifolia Broth., Biblioth. Bot. 87: 9. 1916. Type: Bolivia. Im Hochtal von Viloco, Quimzacruz, ca. 4600 m, Herzog 3159.


Andreaea erythrodictyon Herzog, Beih. Bot. Centralbl. 26(2): 47. 1909. Type: Bolivia. An Steinen un Schneetälchen aufder Nordseite des Cerro Tunari (Prov. Cochabamba), ca. 5100 m, Januar 1908, Herzog s.n.


Andreaea fragilis Müll. Hal., Linnaea 42: 223. 1879. Type: Argentinia. Subtropica alpina, Cuesta de Pinos, inter Grimmiam cespitulum unicum inveni, 27. Majo 1873, Lorentz.

Andreaea karsteniana Müll. Hal., Linnaea 38: 572. 1874. Type: Ecuador. Montis Pichincha, Karsten.


Andreaea laticuspis Broth., Biblioth. Bot. 87: 10. 1916. Type: Bolivia. Huaillattani, Quimzacruz, alt.4900 m, Herzog 2959. Paratype cited: Bolivia. Cord. de Cocapata, Yanakaka, alt. 4500 m, Herzog 4500.


Andreaea lorentziana Müll. Hal., Linnaea 42: 221. 1879. Type: Argentina. Subtropica alpina, Cuesta de la Calderia, 27 Majo 1873, Lorentz. Recorded for Argentina and Bolivia.


Andreaea parallela Müll. Hal., Bot. Jahrb. Syst. 5: 76. 1883. var. gainii (Cardot) Ochyra, Fragm. Florist. Geobot. 41: 479. 1996. Andreaea gainii Cardot, Compt. Rend. Hebd. Séances Acad. Sci. 153: 602. 1911. Type: Antarctica. According to R. Ochyra (pers. comm.) this does not occur in Bolivia.


Andreaea peruviana Broth., Bot. Jahrb. Syst.56 (Beibl.): 123.3. 1920. Type: Peru. Prov. Sandia, oberhalb Ananea, an Felsen, 5100 m, Weberbauer 1045.


Andreaea regularis Müll. Hal., Int. Polarforsch. Deutsch Exped. 2: 286. 1890. Type: ? Reported for Bolivia at 3900 m.


Andreaea robusta Broth., Bot. Jahrb. Syst.49: 174. 1912. Syntypes: Bolivia. Aguila, an der Cordillera Real, 5200 m, Edith Knoche 47, 56.


Andreaea semisquarrosa Müll. Hal., Linnaea 42: 222. 1879. Type: Argentinia. Subtropica alpina, Nevado de Castillo prope Salta, 1873; inter Grimmias crescens, Lorentz.


Andreaea seriata G. Roth, Aussereur. Laubm.42. 3 f. 1. 1911. Type: ? Reported for Ecuador at 5300 m.


Andreaea squarrosa Mitt., J. Linn. Soc., Bot. 12: 629. 1869. Syntypes: Ecuador. Andes Quitenses, Jameson; America australis, Lobb.


Andreaea striata Mitt., J. Linn. Soc., Bot. 12: 628. 1869. Syntypes: Ecuador. Andes Quitenses, Jameson; Brasilia, Sierra dos Orgaos, Gardner 1. Reported for Ecuador (5195 m) and Bolivia (4200-4500 m).


Andreaea tunariensis Broth., Biblioth. Bot. 87: 10. 1 f. 2. 1916. Syntypes: Bolivia. Tunari, alt. ca. 5000 m, Herzog 4812, Herzog 4865.


Andreaea turgescens Schwägr. ex Müll. Hal., Syn. Musc. Frond. 2: 515. 1851. Type: Mexico. In summo m. vulcanico “Pico de Orizaba”, altit. 14000 pedum, Liebmann.


Andreaea vilocensis Broth., Biblioth. Bot. 87: 9. 1916. Syntypes: Bolivia. Hochtal Viloco, Quimzacruz, Herzog 3189, Herzog 3190.


Andreaea vulcanica Lorentz, Moosstudien 156. 1864. Type: Ecuador. Von der höchsten Region des Vulcan’s Ilinitza, 11,000-12,000, Boden Trachyt., 12 Dec 1858, Wagner.


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Plants mostly small, forming dense to loose tufts or mats, dull brown, black to reddish-brown. Stems erect or spreading, several branched; in cross-section cells walls thick-walled, outer 2-3 rows of cells small, inner larger, central strand absent. Leaves ovate-oblong or oblong-lanceolate to ovate-subulate, concave and/or distally channeled above, apex acuminate or narrowly to broadly acute, base subauriculate or not; margins plane, erect or recurved, entire to crenulate by bulging cell walls; ecostate or costate, costa if present short and broad or extending into acumen and subulate; laminal cells thick-walled, smooth or distally papillose on back, upper cells quadrate or short rectangular-rounded; lower and basal cells short to long rectangular, porose or not. Autoicous or dioicous. Perichaetia terminal on stems and branches; leaves differentiated, 2 times or more larger than stem leaves, oblong, convolute. Pseudopodium (seta-like structure) present and ± elongate. Capsules ovate-oblong to obloid, splitting by 4 longitudinal slits or valves, peristome and opercula absent.


Plantas generalmente pequeñas, formando matas o manojos densos a laxos, de color pardo opaco, negro a pardo rojizo. Tallos erectos o patentes, muy ramificados; células con paredes gruesas en corte transversal, 2-3 filas de células externas pequeñas, células internas mayores, sin cordón central. Hojas ovado-oblongas u oblongo-lanceoladas a ovado-subuladas, cóncavas y/o distalmente acanaladas arriba, ápices acuminados o angostamente hasta anchamente agudos, bases subauriculadas o no; márgenes planos, erectos o recurvados, enteros hasta crenulados por abultamientos de las paredes celulares; ecostadas o con costa, costa cuando presente corta y ancha o extendiéndose dentro del acumen y subulada; células de la lámina con paredes gruesas, lisas o distalmente papilosas en el envés, células superiores cuadradas o cortas redondo-rectangulares; células inferiores y basales cortas a largo-rectangulares, porosas o no. Autoicas o dioicas. Periquecios terminales en tallos y ramas; hojas diferenciadas, 2 veces o más grandes que  tallo hojas, oblongas, convolutas. Pseudopodio (estructura como seta) presente y ± alargado. Cápsulas ovado-oblongas hasta obloides, dehiscencia por 4 aperturas longitudinales o valvas, peristoma y opérculo ausentes.


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Keys incomplete…


1. Leaves broadly ovate-oblong to elliptical, apices obtuse-rounded; costa broad and short, 1/2or less lamina length … A. nitida

1. Leaves ovate-lanceolate or -subulate, apices broadly or narrowly acuminate, often bluntly so; costa absent or if present then elongate … 2.

2. Leaves costate; lower lamina cells not porose; perichaetial leaves not papillose … A. subulata

2. Leaves ecostate; lower lamina cells strongly porose; perichaetial leaves papillose... 3.

3. Acumen above expand portion equal in length, narrow and long acuminate … A. acutifolia

3. Acumen above expanded portion short, ca. 1/2 in length, broad and short acuminate or acute … 4.

4. Leaves ovate- to oblong-lanceolate, usually wider at or below mid leaf … A. rupestris

4. Leaves oblanceolate, usually wider above mid leaf … A. brevipes



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