Rather small plants in dense, dull tufts. Stems erect, simple or forked. Leaves contorted when dry, keeled, lanceolate, gradually acuminate, usually acute, sometimes ± awned; margins reflexed to revolute, notched or irregularly serrate near the tip; costa subpercurrent to excurrent, prominent at back; cells subquadrate, thick-walled, and smooth above, short-rectangular below. Dioicous. Perigonia swollen and conspicuous. Perichaetial leaves differentiated, sheathing. Setae erect, elongate; capsules suberect and nearly symmetric to strongly inclined and distinctly asymmetric, ± strumose at the base, ± sulcate when dry; annulus large and revoluble; operculum conic; peristome teeth 16, split nearly to the base into 2 terete divisions connected at the joints toward the base, bordered, papillose. Spores spherical. Calyptrae cucullate, smooth.