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Published In: Flora 12((2) Erganzungsblatter): 19. 1829. (Flora) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 2/17/2011)
Acceptance : Accepted
Project data     (Last Modified On 2/17/2011)

Ptychomitrium has often been associated with the Orthotrichaceae, but its peristome is haplolepidous (Edwards 1979, Crum 1994a) and structurally very close to that of Racomitrium. It has a large, mitrate calyptra that varies from 1/2 to completely covering the capsule and may be plicate or deeply lobed. Calyptrae identical to those of Ptychomitrium also occur in Racomitrium. A distinctive feature of the genus is found in the species with fully bistratose leaves. In those species the leaf cells, especially in the ventral layer, are bulging-mammillose in a manner similar to those of Timmiella anomala. Although Ptychomitrium overlaps with the Grimmiaceae in nearly every feature, the genus is sometimes placed in its own family.


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Ptychomitrium Fürnr., Flora 12 (Erg. 2): 19. 1829. 

Plants small to large in dark-green, blackish or brownish green tufts or cushions. Stems erect, simple or forked. Leaves crisped, inrolled or appressed when dry, erect‑spreading to spreading when wet, oblong to lingulate‑lanceolate or lanceolate; apex cucullate, broadly acute to acuminate; margins erect, recurved or incurved, uni‑ or bi‑stratose, entire or serrate; costa single, strong, subpercurrent to percurrent; lamina uni‑ or bistratose; upper cells rounded‑quadrate, firm‑walled, basal lower cells elongate to oblong or quadrate to short‑rectangular, smooth. Autoicous. Perichaetial leaves not differentiated. Setae single or multiple. Capsules exserted, oblong to cylindrical; opercula long-rostrate; peristome of 16 teeth split nearly to the base into 2–3 papillose divisions. Calyptrae mitrate, deeply lobed, plicate.


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