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Published In: Hedwigia 46(5): 310. 1907. (Hedwigia) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 10/16/2009)
Acceptance : Accepted
Notes     (Last Modified On 10/16/2009)

      Warnstorfia contains 9-10 species. The genus is widespread in mostly cool temperate environments, in the tropical Andes associated with marsh and bogs, shore of ponds and lakes at high elevations. The genus includes species that were previously associated with Calliergon and Drepanocladus.




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Plants medium-sized to large, green, yellowish-green, brown or reddish-brown. Stems few to many branched; in cross-section hyalodermis absent or rarely present (then weakly differentiated and partial), central strand present; pseudoparaphyllia foliose; rhizoid initials inserted at various points on leaf, particularly near leaf apex. Stem leaves straight or falcate from erect to erect-patent base, lanceolate to ovate- or oblong-lanceolate, concave, decurrent or not, apex acuminate or abruptly acute or obtuse-apiculate; margins plane, entire or serrulate (toward apex?); costa single, ca. 2/3 leaf length or long excurrent; median and upper laminal cells linear, smooth, thin- to ± thick-walled, porose or not; alar region transverse-triangular or nearly quadrate in shape, cells inflated, ovoid to oblong, hyaline, mostly thin-walled, occasionally thick-walled; supra-alar cells undifferentiated to differentiated. Branch leaves similar, smaller and narrower. Autoicous or dioicous. Setae elongate. Capsules horizontal, urn cylindrical, curved; opercula conic; peristome double, well developed. Calyptrae cucullate, naked. Spores finely papillose.


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1. Leaf costa short- or long-excurrent in both stem and branch leaves; branch apices pencil-like …. W. trichophylla
1. Leaf costa not excurrent; branch and shoot apices not pencil-like … 2.
2. Stem leaves oblong or ovate, above ± abruptly rounded or narrowing to acute or short-acuminate apex, frequently with short apiculus (best observed in young leaves) … 3.
2. Stem leaves ovate or ovate-lanceolate ovate-triangular base gradually narrowing to acuminate apex, without apiculus … 4.
3. Stem leaf margins entire or at most with one or a few indistinct, obtuse denticles near base of apiculus; leaves ± abruptly narrowing to short apiculus, apex rarely acute or even short-acuminate … W. sarmentosa
3. Stem leaf margins at least partially denticulate, in some leaves distinctly so and frequently coarsely serrulate for a short distance near apex; leaves gradually or abruptly narrowing to acuminate or short-acuminate or sometimes to acute-apiculate or rarely rounded-apiculate apex … W. luipichensis
4. Alar cells forming a large transverse-triangular group, distinctly delimited from surrounding cells; partial hyalodermis usually present; plants often clear red in color … W. exannulata
4. Alar cells forming a rather narrow transverse-triangular group, less distinctly delimited from surrounding cells; hyalodermis absent; plants often reddish-brown in color … W. fluitans
1. Costa foliar corto - o largo-excurrente en hojas de tallos y ramas; ápices de las ramas en forma de lápiz …. W. trichophylla
1. Costa foliar no excurrente; ápice de las ramas e innovaciones no en forma de lápiz … 2.
2. Hojas de los tallos oblongas u ovadas, arriba ± abruptamente redondeadas o estrechamente a ápice agudo o cortamente acuminado, frecuentemente con apículos cortos (mejor observado en hojas jóvenes) … 3.
2. Hojas de los tallos ovadas u ovado-lanceoladas, base ovado triangular estrechándose gradualmente hasta un ápice acuminado, sin apículo … 4.
3. Márgenes de las hojas de los tallos enteras o a lo más con una o unas pocas dentículas indistintas, obtusas cerca de la base del apículo; hojas estrechándose ± abruptamente hasta un apículo corto, ápice raramente agudo o cortamente acuminado … W. sarmentosa
3. Márgenes de las hojas de los tallos al menos parcialmente denticuladas, en algunas hojas distintamente así y frecuentemente ásperamente serruladas por una corta distancia cerca del ápice; hojas estrechándose gradualmente o abruptamente a acuminadas o corto-acuminadas o a veces hasta agudo-apiculadas o raramente con ápice redondeado-apiculado … W. luipichensis
4. Células alares formando un grupo grande, transverso-triangular, claramente delimitado de las células circundantes; hialodermis parcial usualmente presente; plantas generalmente de color rojo claro … W. exannulata
4. Células alares formando un grupo transverso-triangular algo angosto, menos claramente delimitado de las células circundantes; hialodermis ausente; plantas generalmente de color pardo-rojizo … W. fluitans
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