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Published In: Monographs in Systematic Botany from the Missouri Botanical Garden 11: 211. 1985. (Monogr. Syst. Bot. Missouri Bot. Gard.) Name publication detail

Project Name Data (Last Modified On 2/7/2013)
Acceptance : Accepted
Project Data     (Last Modified On 10/8/2017)
Plant Category: Mosses

Notes     (Last Modified On 10/8/2017)
Brymela is distinguished by the rather large habit, the absence of or little developed stem hyalodermis, leaves lacking a border, with leaf margins serrated, of mostly simple teeth, and the ± homogeneous, firm- to thick-walled, porose laminal cells. A neotropical genus of some 12-13 species; in the tropical Andes found in wet lowland to premontane forest, rare at higher elevations; on logs, humus, leaf litter, and rocks. Our species were previously placed in Hookeriopsis.


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Plants small to large and robust, forming loose to dense mats, glossy light or dark green to golden-brown. Stems and branches spreading to subascending; in cross-section central strand absent, hyalodermis absent. Leaves loosely erect-spreading to falcate-secund, oblong to oblong-short lanceolate, 0.8-8.0 mm long, smooth to undulate or rugose, apex acute, short acuminate or subobtuse, base weakly auriculate or not; margins reflexed or recurved in lower 1/3, or at midleaf, subentire to serrulate from about midleaf, often serrate at apex, elimbate or weakly limbate; costae double, 1/2-3/4 leaf length, parallel or slightly diverging, smooth to weakly or strongly serrulate along back; laminal cells smooth, apical cells shorter or not; median cells oblong-linear to linear, smooth to weakly or strongly porose; basal cells short, often golden-brown; marginal cells similar or forming a weak border of long linear cells. Asexual structures, absent. Perichaetia lateral, leaves differentiated, ovate-narrowly lanceolate, sharply serrate. Setae elongate, 10-45 mm long, slender to somewhat stout, smooth to more commonly weakly papillose or roughened below urn. Capsules inclined to horizontal, rarely subpendulous, urn ovoid-short cylindrical, 1-3 mm long, often somewhat flared below mouth when deoperculate; opercula, conic short to long rostrate; peristome, double, exostome teeth strongly furrowed, densely cross-striate, distal tips papillose; endostome basal membrane high, segments keeled and papillose. Calyptrae mitrate-short campanulate, naked, lobed at base. Spores spherical, lightly papillose.
Plantas pequeñas a grandes y robustas, formando tapices ralos a densos, verde claras brillantes o verde oscuras hasta pardo-doradas. Tallos y ramas extendidos a subascendentes; en sección transversal cordón central ausente, hialodermis ausente. Hojas laxamente erecto-extendidas a falcado-secundas, oblongas a oblongo-corto lanceoladas, 0.8-8.0 mm de largo, lisa a ondulada o rugosa, ápice agudo, corto acuminado o subobtuso, base levemente auriculada o no; márgenes reflexos o recurvados en 1/3 inferior, o en la mitad de la hoja, subenteros a serrulados desde cerca de la mitad de la hoja, generalmente aserrados en el ápice, elimbados o levemente limbados; costa doble, 1/2-3/4 del largo de la hoja, paralelas o ligeramente divergentes, lisas a levemente a fuertemente serruladas a lo largo del dorso; células de la lámina lisas, células apicales más cortas o no; células mediales oblongo-lineares a lineares, lisas a levemente o fuertemente porosas; células basales cortas, generalmente pardo-doradas; células marginales similares o formando un borde débil de células lineares largas. Estructuras asexuales, ausentes. Periquecios laterales, hojas diferenciadas, ovado-angostamente lanceoladas, fuertemente aserradas. Setas alargadas, 10-45 mm de largo, delgadas a algo gruesas, lisas a más comúnmente levemente papilosas o rugosas debajo de la urna. Cápsulas inclinadas a horizontales, raramente subpéndulas, urna ovoide-corto cilíndrica, 1-3 mm de largo, generalmente algo acampanada debajo de la boca luego de desoperculada; opérculo, cónico corto a largo rostrado; peristoma doble, dientes del exostoma fuertemente surcados, densamente estriados transversalmente, extremos distales papilosos; membrana basal del endostoma alta, segmentos plegados y papilosos. Caliptra mitrada-corta campanulada, desnuda, lobada en la base. Esporas esféricas, ligeramente papilosas.

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1. Leaf apices acute to obtuse-rounded … 2.
1. Leaf apices acuminate …4.
2. Leaves falcate to falcate-secund, mostly less than 2 mm long… B. obtusifolia
2. Leaves loosely erect to erect-spreading, ± straight, not falcate, leaves mostly greater than 2 mm long … 3.
3. Leaf margins sharply serrate, teeth strongly bifid and swollen … B. rugulosa
3. Leaf margins weakly and bluntly serrulate, few teeth bifid but not swollen … B. parkeriana
4. Leaf apices cuspidate, with a notched sinuse at junction with lamina and leaf tip … B. cuspidata (and related forms)
4. Leaf apices gradually to± abruptly acuminate, neither cuspidate nor notched … 5.
5. Leaves 3.5-4 mm long … B. “antioquiana”
5. Leaves 2.4 mm long or less … 6.
6. Leaf apices short acuminate … B. acuminata
6. Leaf apices long and slenderly acuminate …  B. websteri
1. Apices foliares agudos a obtuso-redondeados … 2.
1. Apices foliares acuminados … 4.
2. Hojas falcadas a falcado-secundas, mayormente menores de 2 mm de largo … B. obtusifolia
2. Hojas laxamente erectas a erecto-extendidas, ± rectas, no falcadas, hojas mayormente mayores de 2 mm de largo … 3.
3. Márgenes foliares agudamente aserrados, dientes fuertemente bífidos e hinchados … B. rugulosa
3. Márgenes foliares levemente y obtusamente serrulados, con pocos dientes bífidos, pero no inflados … B. parkeriana
4. Apices foliares cuspidados, con un seno hendido en la unión entre la lámina y el ápice foliar … B. cuspidata (y formas relacionadas)
4. Apices foliares gradualmente a ± abruptamente acuminados, no cuspidados ni hendidos … 5.
5. Hojas 3.5-4 mm de largo … B. “antioquiana”
5. Hojas 2.4 mm de largo o menos … 6.
6. Apices foliares corto acuminados … B. acuminata
6. Apices foliares largos y delgadamente acuminados …  B. websteri
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