Plants yellow-green above, brown below, irregularly branched, weakly radiculose, up to 3 cm high. Leaves incurved-contorted and occasionally catenulate when dry, spreading up to 45¡ when moist, usually very fragile, weakly concave, commonly 3-5 mm long, long-lanceolate; costa with ventral epidermal cells quadrate and papillose except toward the apex where they are elongate and smooth; upper cells 10-12(-14) µm, usually evenly thick-walled, with papillae 2-4 and bifid or sometimes fused to form a single compound papilla covering the lumen; basal cells with thin or thick walls, 10-15 µm wide and 3-5:1, generally short-rectangular and thick-walled near the top of the sheathing base. Dioicous. Setae 1-1.5 cm long; capsules 3-3.2 mm long, long-cylindric; annulus of 1-2 rows of vesiculose cells; operculum long-conic to rostrate; peristome teeth long-ligulate, up to 220 µm long, yellow to brown, weakly papillose to striolate. Spores 13-20 µm, papillose.