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Published In: Die Laubmoose Deutschlands, Oesterreichs und der Schweiz 1: 592. 1888. (Laubm. Deutschl.) Name publication detail

Project Name Data (Last Modified On 10/22/2013)
Acceptance : Accepted
Project data     (Last Modified On 10/22/2013)
Nomenclature: T. anomala (B.S.G.) Limpr., Laubm. Deutschl. 1: 592. 1888.
Barbula anomala B.S.G., Bryol. Eur. 2(fasc. 13/15). 1842.
Trichostomum subanomalum Besch., Mem. Soc. Sci. Nat. Cherbourg 16: 177. 1872.
Timmiella subanomala (Besch.) Broth. in E. & P., Nat. Pfl. 1(3): 396. 1902.
Distribution: On calcareous soil, rock, or bases of trees at moderate to high altitudes; Baja California, Baja California Sur, Chihuahua, Coahuila, Distrito Federal, Durango, Guerrero, Hidalgo, Jalisco, Mexico, Michoac‡n, Morelos, Oaxaca, Puebla, Queretaro, San Luis Potos’, Sinaloa, Sonora, Tamaulipas, Tlaxcala, Veracruz. - Mexico; Guatemala and El Salvador; southwestern United States; southern Europe; Himalayas.
Discussion: A Mexican report of a sterile collection of T. crassinervis (Hampe) L. Koch needs verification (Bowers, Delgadillo & Sharp, 1976). The monoicous sexuality of T. anomala provides the only certain means of distinguishing it from that species of western North America.
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Plants 15-20 mm high. Stems with a strong central strand (and occasionally hollow) and a hyaloderm; axillary hairs hyaline. Leaves up to 4 or 5 mm long, somewhat broadened above a short, erect base; margins bluntly and remotely serrate above the middle, serrulate below (usually nearly to the base); costa percurrent, covered ventrally by short, bulging-mammillose cells and dorsally by elongate, smooth cells, with 10-16 guide cells in 1 layer and hydroid strands (Begleiter cells) below and sometimes above them and 3-5 parenchymatous cells between the ventral stereids and the epidermis; upper cells 9-12 µm; basal cells 10-18 µm wide. Autoicous (reported to be synoicous as well, perhaps also rhizautoicous). Setae 15-20 mm long, yellow- to red-brown, clockwise-twisted; capsules 3-5 mm long, brown, ± striolate when dry; annulus up to 5 cell layers thick, revoluble or deciduous in fragments; operculum about 1.3 mm high, high-conic, with cells in straight or weakly clockwise-twisted rows; exothecial cells elongate; peristome teeth 300-650 µm long, straight or weakly twisted in a clockwise fashion. Spores 10-13 µm, yellowish, weakly papillose.


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