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Published In: Bericht über die Thätigkeit der St. Gallischen Naturwissenschaftlichen Gesellschaft 1873–74: 117 (Gen. Sp. Musc. 1: 579). 1875. (Ber. Thätigk. St. Gallischen Naturwiss. Ges.) Name publication detail
 

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Discussion:

Brachymenium systylium is a small species with usually erect-imbricate leaves. The leaves are long and narrow with subpercurrent costae toward the base but ovate to oval with excurrent costae near the apical parts of the stem. It differs from most other regional species of Brachymenium in having upper leaves with long, entire or serrate, capillary hair-points. There are two other Central American species of Brachymenium with the gametophytic aspect of B. systylium: B. niveum and B. morasicum. In  B. niveum the leaf hair-points are entirely hyaline, and its uppermost leaf cells are narrowly elongate. In contrast the hair-points of B. systylium are reddish at base, and its upper leaf cells are usually rhomboidal-hexagonal. As noted by (Ochi 1994) the type of B. condensatum (a synonym of B. systylium) has upper leaf cells intermediate between B. systylium and B. niveum. Brachymenium morasicum differs from B. systylium in having toothed to sharply serrate or denticulate upper leaf margins, well-developed leaf limbidia, and long, narrowly cylindrical capsules.

Illustrations: Duby (1870, Pl. 1 4, as Bryum angolense); Fleischer (1904, Fig. 99, as Bryum garutense); Williams (1923, Pl. 1 1–14); Brotherus (1924, Fig. 124, as Brachymenium crassinervium); Andrews (1935, Fig. 82A); Bartram (1949, Fig. 73 E–G); Ochi (1964, Fig. 14, as B. leptostomoides); Ochi (1972, Fig. 7, as B. angolense); Crum and Anderson (1981, Fig. 232); Ochi (1985, Fig. 3 F–M); Shaw (1985a, Pl. 5 26–28); Magill (1987, Figs. 9–15, as B. angolense); Sharp et al. (1994, Fig. 375 e–i). Figure 137 A–C.
Habitat: On tree trunks and branches (Quercus, Pinus), logs, rocks and boulders, or wet soil; 950–3000 m.
Distribution in Central America: GUATEMALA. Alta Verapaz: Standley 69650 (F); Baja Verapaz: Sharp 2892 (FH); Chimaltenango: Standley 61856 (F, FH); Huehuetenango: Standley 82867 (F); Quezaltenango: Standley 85738 (F); Quiché: Hermann 26367 (F); Sacatepéquez: Standley 59011 (F, FH, MICH); San Marcos: Smith et al 934a (F); Sololá: Steyermark 46959 (F, FH, MICH, MO); Totonicapán: Molina et al. 30423 (F). HONDURAS. Comayagua: Allen 14031 (MO, TEFH); Intibucá: Standley 25573 (F, MICH); La Paz: Allen 14549 (MO, TEFH); Lempira: Allen 11195 (MO, TEFH); Francisco Morazán: Standley 26402 (F, MICH); Olancho: Allen 12470 (MO, TEFH). NICARAGUA. Estelí: Stevens 15020a (MO); Matagalpa: Richards et al. 3674 (F, MICH, MO). COSTA RICA. Alajuela: Standley 34657 (FH); Cartago: Alfaro 114 (F); Heredia: Crosby 3878 (MICH, MO); Puntarenas: Alfaro (FH); San José: Griffin & Morales B89 (F, MO). PANAMA. Chiriquí: Allen 9061 (MICH, MO, PMA).
World Range: Southwestern, South-Central, and Southeastern U.S.A.; Mexico; Central America; Western South America, Brazil; West-Central,  East, and South Tropical Africa, Southern Africa; China; Indian Subcontinent, Indo-China, Malesia.

 

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Brachymenium systylium (C. Müll.) Jaeg., Ber. Thätigk. St. Gallischen Naturwiss. Ges. 1873–74: 117. 1875.

Bryum systylium C. Müll., Syn. Musc. Frond. 1: 320. 1848. Protologue: Mexico. Prope Xalapam, Deppe & Schiede.

Brachymenium brachypelma C. Müll. ex Ren. & Card., Bull. Soc. Roy. Bot. Beligique 31(1): 162. 1893. Protologue: Costa Rica. Chemin de la Piedra Blanca, Pittier 5562; Forêts du Barba, Pittier 5563.

Brachymenium condensatum Williams, Bryologist 26: 2. 1923. Protologue: Mexico. Michoacan, Patzcuaro, Dec. 21, 1891, Pringle 762 (MO).

Plants small, bright-green to yellowish green, in dense, shiny tufts or mats. Stems 5-30 mm long, epidermal cells rectangular, densely tomentose at base; rhizoids reddish brown, papillose, at times with red-brown, multicellular, globose propagula. Leaves crowded in rosulate tufts, 1.2–2.5 mm long, erect-appressed to imbricate not or rarely somewhat spirally twisted, spreading when wet, concave, ovate, oval, oblong to oblong-lanceolate; apices short acuminate, ending in a smooth or toothed capillary hair-point, yellowish at base, hyaline in upper ½, 0.4–1.0 mm μm long; margins not or weakly bordered below by 1-2 rows of somewhat narrower, elongate cells, upper marginal cells short-rectangular to somewhat elongate, entire or weakly serrulate, plane above, narrowly recurved below; costa ending below the apex in lower leaves, stoutly long-excurrent in upper leaves, ventral superficial cells at base quadrate to rectangular; cells firm-walled throughout, upper cells hexagonal to rhomboidal-hexagonal, 30–60 x 12 μm, basal cells often reddish near leaf insertion, quadrate to short-rectangular in up to 20 cells along the basal margin, 20–30 x 20 μm. Dioicous. Setae 15–25 mm long, yellowish red. Capsules 2.5–4.0 mm long, obovate-oblong, constricted at neck, erect to suberect; opercula 0.3–0.5 mm long, conic-apiculate; annuli compound, revoluble; exostome teeth reddish yellow, densely papillose, endostome yellowish, papillose, basal membrane to ½ the exostome teeth length, segments rudimentary, cilia rudimentary to absent. Spores 18–24 μm, papillose. Calyptra not seen.

 

 

 
 
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