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Published In: Flora 12: 393, pl. 3. 1829. (Flora) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 10/22/2013)
Acceptance : Accepted
Project data     (Last Modified On 10/22/2013)
Nomenclature: T. brachydontium Bruch ex F. Muell., Flora 12: 393. 1829.
T. mutabile Bruch ex De Not., Syll. 192. 1838.
Symblepharis jamaicensis C. Muell., Bull. Herb. Boissier 5: 555. 1897.
Hyophila mexicana Ther., Smithson. Misc. Coll. 85(4): 13. 1931.
Didymodon planifolius P.-Varde & Ther., Rev. Bryol. Lichenol. 14: 11. 1944.
Distribution: On soil, sometimes on roadbanks, 300-2500 m alt.; Baja California, Chiapas, Chihuahua, Coahuila, Colima, Distrito Federal, Durango, Hidalgo, Jalisco, Mexico, Nuevo Leon, Nayarit, Oaxaca, Puebla, San Luis Potos’, Sonora, Tamaulipas, Tlaxcala. - Mexico; Central and South America; West Indies; Texas; Europe, Africa, Asia, Australasia.
Discussion: Robust plants resemble T. crispulum in having erect leaf margins and costa darker than the lamina (in mature leaves), but the stout, recurved mucro and leaf base not much differentiated in shape are diagnostic. The peristome is variable in development, and therefore certain South American plants lacking a peristome can probably be referred to T. brachydontium. (Such plants might be confused with Hyophila.) Small plants can generally be distinguished from Weissiae by plane or erect leaf margins, in comnbination with a stout mucro (but see W. sinaloensis) and from Barbula convoluta by the smooth dorsal surface of the costa and deep yellow-orange color in 2% KOH solution (as opposed to B. convoluta, which has a papillose costa and stains light yellow in KOH).
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Plants green above, brown below, up to 2.5 cm high. Leaves crowded, tubulose and incurved when dry, spreading up to 70¡ when moist, nearly flat to broadly concave, 2-2.5(-3) mm long, ligulate to lanceolate, rounded to sharply acute at the apex; margins plane to erect; costa often very thick, short-excurrent as a stout mucro; upper cells subquadrate to hexagonal or short-rectangular, 6-9 µm wide and 1(-2):1, thin- to evenly thick-walled, each with 2-4 bifid to multifid papillae; basal cells differentiated across the insertion, rectangular, 9-13 µm wide and 2-4:1, thin-walled, hyaline. Dioicous. Setae 3.5-10 mm long; capsules 0.8-1.5 mm long, cylindric to broadly ellipsoid; annulus of 1-2 rows of usually weakly vesiculose cells, persistent, fragmenting, or revoluble; operculum 0.5-0.8 mm long, conic; peristome teeth often rudimentary, 30-300 µm long, straight, ligulate to subulate, entire or 2-3-forked, yellow to red-brown, spiculose to minutely reticulate, with basal membrane short or lacking. Spores 15-20 µm, strongly papillose.


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