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Published In: Muscologiae Recentiorum Supplementum 4: 79. 1819[1818]. (Muscol. Recent. Suppl.) Name publication detail

Project Name Data (Last Modified On 3/16/2009)
Acceptance : Accepted
Project Data     (Last Modified On 3/16/2009)
General Reference:

Illustration & Map     (Last Modified On 3/16/2009)
Related Objects:
Illustrations: X.-J. Li (ed.) 2000 (Pl. 26, figs. 12–20).

Click here for distribution map.

Map: Racomitrium heterostichum (Hedw.) Brid. (Distribution in China)

Distribution     (Last Modified On 3/16/2009)
Habitat: on rocks from lowlands to high mountains.
Distribution: China, Japan, Europe, northern Africa, and North America.

Specimens Examined     (Last Modified On 3/16/2009)
Specimens Examined:
Chinese specimens examined: Hubei: Mt. Shennongjia, 1980 Sino-Amer. Exped. Team 1139e (MO, PE). Jilin: Mt. Changbai, C. Gao 22248 (IFSBH, MO). Shaanxi: Mt. Qinling, Q. Huang 1298 (MO, PE). Sichuan: Ba-tang Co., S. He 31616 (MO, PE); Xiang-cheng Co., S. He 31759 (MO, PE). Taiwan: Hua-lien Co., C.-C. Chuang 5567 (MO, UBC), Shevock 14426 (CAS, MO); Ilan Co., C.-C. Chuang 1862 (MO, UBC); Miao-li Co., Shevock 18085 (CAS, MO); Nan-tou Co., Shevock 17951 (CAS, MO); Ping-tung Co., C.-C. Chuang 1504 (MO, UBC); Tai-chung Co., Shevock 18073 (CAS, MO).

Notes     (Last Modified On 3/16/2009)
general taxon notes:
Racomitrium heterostichum is characterized by having stems densely branched, leaves often secund, long, dentate, hyaline hair-points, short and smooth median leaf cells, and scarcely differentiated alar cells. Crum and Anderson (1981) described Racomitrium heterostichum as having bistratose leaf cells at the margins with the exception of R. heterostichum var. microcarpum, which has unistratose leaf cells at the margins. In Asia, Racomitrium heterostichum usually has unistratose leaf cells at the margins except for R. heterostichum var. brachypodium (Besch.) Nog. and R. heterostichum var. sudeticum (Funck) Bauer. We do not recognize these two varieties under R. heterostichum.


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12. Racomitrium heterostichum (Hedw.) Brid.   异枝砂藓   yi-zhi sha xian
Muscol. Recent. Suppl. 4: 79. 1819[1818]. Grimmia heterosticha (Hedw.) C. Müll., Syn. Musc. Frond. 1: 807. 1849. Trichostomum heterostichum Hedw., Sp. Musc. Frond. 109. 1801. Type. Germany.

Plants usually large to robust, up to 8–10 cm long, yellowish green above, dark brownish or blackish brown below, often appearing grayish due to hyaline hair-points, in dense cushions. Stems densely branched with long or short branches; central strand absent. Leaves imbricately appressed when dry, erect-spreading when moist, narrowly ovate-lanceolate, 2.5–3.5 mm long, keeled above, sometimes secund, with long, dentate, hyaline hair-points, the hair-points often decurrent at base; margins recurved on both sides; costa percurrent; upper and median leaf cells quadrate to short-rectangular, 10–20 µm × 8–10 µm, sinuosely thick-walled, smooth, unistratose at margins; basal cells rectangular to elongate-rectangular, 15–35 µm × ca. 10 µm, strongly sinuosely thick-walled; alar cells scarcely differentiated, with only a row of 3–10 slightly larger, rectangular, straight-walled cells. Perichaetial leaves smaller than stem leaves, ovate-lanceolate, narrowly obtuse at apex, with shorter hyaline hair-points. Setae erect, 4–8 mm long; capsules cylindrical, 1.5–3.0 mm long; opercula long-rostrate; peristome teeth linear-lanceolate, divided to the base, densely papillose. Spores spherical, 14–17 µm in diameter.



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