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Published In: Akademie der Wissenschaften in Wien, Sitzungsberichte, Mathematisch-naturwissenschaftliche Klasse, Abteilung 1 133: 567. 1924. (Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1,) Name publication detail

Project Name Data (Last Modified On 3/16/2009)
Acceptance : Accepted
Project Data     (Last Modified On 5/4/2009)
General Reference:

Illustration & Map     (Last Modified On 5/4/2009)
Illustrations: Grimmia handelii Broth. (Figs. 1–12.)
Map: Grimmia handelii Broth. (Distribution in China)

Distribution     (Last Modified On 5/4/2009)
Habitat: on rocks in alpine regions; alt. 4300–4500 m.
Distribution: endemic to China.

Specimens Examined     (Last Modified On 5/4/2009)
Specimens Examined:
Chinese specimens examined: Sichuan: Xiang-cheng Co., M.-Z. Wang 813836 (PE). Yunnan: Li-jiang Co., Mt. Yulong, W.-X. Xu 0011-1 (HKAS)

Notes     (Last Modified On 5/4/2009)
general taxon notes:
Grimmia handelii is characterized by having muticous, keeled leaves, rather small upper and median leaf cells, and slender, filiform young branches originating from the bases of older stems. Grimmia handelii is close to G. unicolor in having similar muticous leaves, but is distinguished from the latter by the keeled leaves with acute apices. In constrast, Grimmia unicolor has somewhat cucullate leaves with obtusely rounded apices.


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9. Grimmia handelii Broth.   韩氏紫萼藓   han-shi zi-e xian
Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1, 133: 567. 1924. Type. China: southwestern Sichuan (Setschwan austro-occid.), Yuan-yan (Yenyuen) Co., 28°15′N, alt. 4150–4300 m, Handel-Mazzetti 2667 (holotype H; isotype S).
Plants up to 4 cm high, in loose tufts, yellowish green above, dark brown to blackish below. Stems sparsely branched, easily separated from each other, with developed central strand and bulging outer surface of epidermis in cross section; filiform young shoots often arising from basal part of older stems, leaves of the shoots small, closely appressed, rounded-ovate, mucronate, 0.30–0.36 mm × 0.25–0.34 mm. Branch leaves imbricate, slightly contorted when dry, erect-spreading when moist, 1.7–2.2 mm long, lanceolate, keeled above, muticous, acute, without hyaline hair-points; margins narrowly recurved on one side; costa strong, single, ending just below the apex, with several small median cells in cross section; upper and median cells unistratose except at margins, small, ca. 5 µm wide, irregularly quadrate, thick-walled, somewhat sinuose; basal marginal cells hyaline, rectangular, 23–46 µm × 7–9 µm, straight, thin-walled; basal juxtacostal cells elongate, 35–46 µm × 5–7 µm, with straight walls. Dioicous. Sporophytes and calyptrae not seen.
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