9. Grimmia handelii Broth. 韩氏紫萼藓 han-shi zi-e xian
Akad. Wiss. Wien Sitzungsber., Math.-Naturwiss. Kl., Abt. 1, 133: 567. 1924. Type. China: southwestern Sichuan (Setschwan austro-occid.), Yuan-yan (Yenyuen) Co., 28°15′N, alt. 4150–4300 m, Handel-Mazzetti 2667 (holotype H; isotype S).
Plants up to 4 cm high, in loose tufts, yellowish green above, dark brown to blackish below. Stems sparsely branched, easily separated from each other, with developed central strand and bulging outer surface of epidermis in cross section; filiform young shoots often arising from basal part of older stems, leaves of the shoots small, closely appressed, rounded-ovate, mucronate, 0.30–0.36 mm × 0.25–0.34 mm. Branch leaves imbricate, slightly contorted when dry, erect-spreading when moist, 1.7–2.2 mm long, lanceolate, keeled above, muticous, acute, without hyaline hair-points; margins narrowly recurved on one side; costa strong, single, ending just below the apex, with several small median cells in cross section; upper and median cells unistratose except at margins, small, ca. 5 µm wide, irregularly quadrate, thick-walled, somewhat sinuose; basal marginal cells hyaline, rectangular, 23–46 µm × 7–9 µm, straight, thin-walled; basal juxtacostal cells elongate, 35–46 µm × 5–7 µm, with straight walls. Dioicous. Sporophytes and calyptrae not seen.