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Published In: English Botany 24: 1728. 1807. (Engl. Bot.) Name publication detail

Project Name Data (Last Modified On 3/16/2009)
Acceptance : Accepted
Project Data     (Last Modified On 3/16/2009)
General Reference:

Illustration & Map     (Last Modified On 3/16/2009)
Related Objects:
Illustrations: X.-J. Li (ed.) 2000 (Pl. 12, figs. 1–10); T. Cao & Vitt 1986 (Fig. 38).

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Map: Grimmia pulvinata (Hedw.) Sm. (Distribution in China)

Distribution     (Last Modified On 3/16/2009)
Habitat: on exposed rocks or cliff faces.
Distribution: China, India, Pakistan, Turkmenistan, Turkey, Ukraine, Russia, Europe, Africa, North and South America, Australia, and New Zealand.

Specimens Examined     (Last Modified On 3/16/2009)
Specimens Examined:
Chinese specimens examined: Taiwan: Hua-lien Co., Koponen 18110 (ALTA, H). Xizang: Long-zi Co., M. Zang 939, 1362, 1652 (all in HKAS); Lang-ka-zi Co., Xizang Exped. Team 7594 (PE); Qu-shui Co., M. Zang 22 (HKAS).

Notes     (Last Modified On 3/16/2009)
general taxon notes:
Grimmia pulvinata is characterized by its small, dense, hoary cushion habit and hyaline, long-awned leaf apices. The hair-points are slender and nearly smooth, usually at least half to the full leaf length. The leaf cell walls are straight or nearly so. The capsules are distinctly wrinkled-striate when dry and positioned on long, curved setae. The sexual condition is autoicous, with perigonia located just below the perichaetia. The present species is closely related to G. fuscolutea. Both taxa have curved setae, straight basal leaf cell walls, and an autoicous sexual condition. Grimmia pulvinata differs from G. fuscolutea in having broader leaves with much longer awns, sometimes as long as the lamina length and with straight or slightly sinuose upper and median leaf cell walls. C. Gao (1977) and Koponen et al. (1983) reported this species from Mt. Changbai; however, the specimens were misidentified and belong to G. fuscolutea.


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20. Grimmia pulvinata (Hedw.) Sm.   垫丛紫萼藓   dian-cong zi-e xian
Engl. Bot. 24: 1728. 1807. Dryptodon pulvinatus (Hedw.) Brid., Bryol. Univ. 1(1): 196. 1826. Campylopus pulvinatus (Hedw.) Brid., Muscol. Recent. Suppl. 4: 75. 1819[1818]. Trichostomum pulvinatum (Hedw.) Web. & Mohr, Bot. Taschenbuch. 109. 1807. Dicranum pulvinatum (Hedw.) Lag., D. García & Clemente, Anales Ci. Nat. 5(14): 176. 1802. Fissidens pulvinatus var. communis Hedw., Sp.. Musc. Frond. 158. 1801, nom. illeg. incl. typ. spec. Fissidens pulvinatus Hedw., Sp. Musc. Frond. 158. 1801. Type. Germany: “in tectis et muris vulgaris, vere malurans” (lectotype G, designated by T. Cao & Vitt 1986).
Plants up to 1.5–2.0 cm high, yellowish green above, brownish below, in small, dense, hoary cushions. Stems moderately branched, with well developed central strand in cross section. Leaves imbricate, erect when dry, spreading when moist, 2.2–3.1 mm long, elliptic- to oblong-lanceolate, obtusely keeled above, ending in a long, slender, faintly toothed hyaline hair-point, up to half to full length of leaf; margins narrowly recurved on one or both sides in the lower part, thickened and bistratose toward apex; costa percurrent, consisting of small abaxial cells in cross section; upper cells more or less unistratose except at margins, rounded-quadrate, with incrassate and slightly sinuose walls, 5–7 µm wide; median cells quadrate to short-rectangular, 9–11 µm × 6–8 µm, with nearly straight walls; basal cells short-rectangular to rectangular, almost uniform in size, straight and thin-walled, 11–40 µm × 7–9 µm. Autoicous. Perichaetial leaves larger than upper stem leaves, with elongate hyaline hair-points; perigonia located just below perichaetia. Setae long, distinctly curved when moist; capsules horizontal or pendent, elliptic, distinctly 8-striate when dry; exothecial cells rounded-hexagonal to elongate-hexagonal, thin-walled; stomata present near base of urn; annuli consisting of well developed elongate cells with thick walls; opercula with short, straight or somewhat oblique beaks; peristome teeth erect, lanceolate, yellowish brown to reddish brown, imperfectly cleft in 2–3 branches and perforate above; both outer and inner surfaces densely papillose above, papillose below. Calyptrae mitrate. Spores 10–12 µm in diameter, minutely granulose.


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