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Published In: Hedwigia 46(5): 309. 1907. (Hedwigia) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 2/27/2009)
Acceptance : Accepted
Project Data     (Last Modified On 5/5/2009)
General Reference:

Illustration & Map     (Last Modified On 5/5/2009)
Illustrations: Sanionia uncinata (Hedw.) Loeske (Figs. 1–10.)
Map: Sanionia uncinata (Hedw.) Loeske (Distribution in China)

Distribution     (Last Modified On 5/5/2009)


Habitat: on soil, rocks, decayed wood, and tree trunks, usually in dry coniferous forest; alt. 490–4700 m.

Distribution: China, Japan, India, Nepal, Pakistan, Greenland, Europe, Russia, Africa, America, Mexico, Oceania, and Antarctica.

Specimens Examined     (Last Modified On 5/5/2009)
Specimens Examined:


Chinese specimens examined: Hebei: Wei Co., C. Gao 30584, 31034 (both in IFSBH); Xing-long Co., J.-X. Ji 376 (MO); Zhang-bei Co., Q.-W. Wang 2006 (PE); Zhuo-lu Co., Q.-W. Wang 1438, 1681 (both in PE). Heilongjiang: Hua-chuan Co., X.-G. Li 73 (PE); Ning-an Co., X.-L. Zhu 217 (PE), C. Gao & M.-X. Nan 6549 (IFSBH); Xiao Xing An Ling (Mt.), X.-L. Zhu 162 (PE), Y.-H. Wu 1240 (IFSBH); Yichun City, C. Gao 239, 262 (both in IFSBH). Hubei: Site unknown, Y.-S. Zou 2-0030 (IFSBH). Jilin: Chang-bai Co., C. Gao 7306, 7582 (both in IFSBH); Jiao-he Co., X.-G. Kang 1623, C. Gao 1634 (both in IFSBH); Mt. Changbai, C. Gao 1192 (MO), Koponen 36829 (IFSBH, PE), Y.-H. Wu 988 (IFSBH); Wang-qing Co., C. Gao & G.-C. Chang 8653, 8912 (both in IFSBH). Liaoning: Kuan-dian Co., T. Cao & S.-Z. Wang 38636 (MO), Y.-H. Wu 0919 (IFSBH), Buck 23788 (NY). Nei Mongol: Aershan City, C. Gao 3264 (IFSBH, MO); Gen-he, H.-F. Xie 060, C. Gao 812 (both in IFSBH); Mt. Helan, Z.-G. Tong 1361 (MO, PE); Wufeng, C. Gao 865 (MO); Zhalan-tun Co., X.-L. Bai 2653 (NY). Qinghai: Hu-zhu Co., M. Zang 6 (HKAS), Long 27139 (E). Shaanxi: Hu Co., Z.-P. Wei 4798 (PE); Mt. Qinling, P.-C. Chen et al. 514 (PE); Mt. Taibai, Giraldi 992 (as Drepanocladus sinensi-uncinatus, F), Giraldi 885 (as Drepanocladus filicalyx, F, NY), 996 (as Drepanocladus sinensi-uncinatus, BM, F), 2093 (NY). Shanxi: Ning-wu Co., C. Gao 30871 (IFSBH, MO). Sichuan: A-ba Co., S. He 31008 (MO); Bao-xing Co., Q. Li 3300 (IFSBH); Cuo-si-jia Co., Z.-T. Guan 5468, Exped. Team 3015 (both in PE); Da-jin Co., Z.-T. Guan 3753, 4931 (both in PE); Dan-ba Co., Anonymous 5574 (PE); Dao-cheng Co., M.-Z. Wang 814072 (PE); Dao-fu Co., Z.-T. Guan 11022, 17-0310 (both in PE); De-ge Co., J.-L. Xiang s.n., 2034 (both in PE); Li Co., Allen 6798 (MO); Lu-huo Co., Anonymous 7203101 (PE); Ma-er-kang, S. He 31234 (MO), Q. Li 3530, X.-J. Li 1312 (both in HKAS); Mao-wen Co., Q.-Z. Zheng et al. 141, Q. Li 06884 (both in IFSBH); Nan-ping Co., S. He 30130a (MO, PE); Song-pan Co., S. He 30542a (MO, NY); Tian-quan Co., C. Gao et al. 17685, Q. Li 4768 (both in IFSBH); Wo-long Co., Q.-Z. Zhen 141 (MO); Xi-chang Co. (Ning-yuen), Handel-Mazzetti 336 (E); Xiang-cheng Co., Exped. Team 4391 (PE); Xiao-jin Co., Exped. Team 4931, J.-S. Zhao 624 (both in HKAS); Ya-jiang Co., S. He 31935 (MO). Taiwan: Hua-lien Co., S. He 36004 (MO); Nan-tou Co., S. He 36181 (MO). Xinjiang: Bur-jin Co., S.-L. Chen 313, Q.-T. Jiang 316 (both in PE); Di-hua Co., S.-E. Liu 3034 (MO); Fu-hai Co., B.-Q. Li 023, S.-L. Chen 34 (both in PE); Fu-kang Co., J.-C. Zhao & R.-L. Dang 03246, 3277 (both in IFSBH); Fu-yun Co., R.-C. Qin 2004, B.-Q. Li 654 (both in PE); Ha-mi Co., S.-L. Chen 19 (PE); He-jing Co., S. He 951199 (MO); Mt. Altai, S.-L. Chen 2, R.-C. Qin 3129 (both in PE), C.-Y. Yang 436 (MO); Ni-le-ke Co., J.-C. Zhao 03766 (IFSBH); Qi-tai Co., B. C. Tan 93-1055 (E); Ürümqi City, S.-E. Liu 3034 (PE), J.-C. Zhao 953129 (MO); Yi-li Co., S.-L. Chen 30, 61 (both in PE); Zhao-su Co., J.-C. Zhao 4202 (IFSBH). Xizang: Ba-qing Co., K.-Y. Zhao 44 (PE); Cha-yu Co., Q.-W. Wang 6639, M.-Z. Wang 8518 (both in PE); Ding-jie Co., K.-Y. Lang 1157a (MO, PE); Ha-ba-he Co., Y.-R. Lin 12 (PE); Mt. Everest, X.-G. Wang 99 (PE); Nie-la-mu Co., Exped. Team 7569 (PE); Ri-dong Co., M. Zang 6363, 6406 (both in HKAS); Ya-dong Co., Exped. Team 7860, 7878, 7881 (all in PE), M. Zang 795 (HKAS). Yunnan: Da-li Co., Redfearn et al. 1396 (MO); Di-qing Co., Long 24030, 24082 (both in E, S); Gong-shan Co., Shevock 23331 (CAS, MO), D.-C. Zhang 886 (HKAS); Mt. Yulong, Long 18833, 19000 (both in E, S); Shangri-la Co., Handel-Mazzetti 1224 (E); Wei-xi Co., W.-C. Hu 10 (PE), Long 24556 (E, S).

Notes     (Last Modified On 5/5/2009)
general taxon notes:


Sanionia uncinata usually grows in the dry and terrestrial environments. It is easily recognizable in the field by its strongly falcate-secund leaves with a strong, single costa.


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1. Sanionia uncinata (Hedw.) Loeske   三洋藓   san yang xian
Hedwigia 46: 309. 1907. Drepanocladus uncinatus (Hedw.) Warnst., Beih. Bot. Centralbl. 13: 417. 1903. Hypnum uncinatum Hedw., Sp. Musc. Frond. 289. 1801. Type: Europe.
Drepanocladus uncinatus fo. auriculatus Mönk., Laubm. Eur. 788. 1927.
Drepanocladus uncinatus fo. longicuspis Smirnova, Not. Syst. Cryptog. Inst. Bot. Kom. Act. Sci. URSS 9: 197. 1953.
Drepanocladus uncinatus fo. plumulosus Mönk., Laubm. Eur. 788. 1927.
Hypnum filicalyx (Müll. Hal.) Paris, Index Bryol. Suppl. 200. 1900, nom. illeg. Drepanocladus filicalyx Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 123. 1896, nom. illeg. Type: China. Shaanxi (Schen-si) Kuan-tou-san, Giraldi 885 (lectotype F, isotype NY).
Hypnum sinensi-uncinatum Paris, Index Bryol. Suppl. 212. 1900. Drepanocladus sinensi-uncinatus Müll. Hal., Nuovo Giorn. Bot. Ital., n.s., 3: 123. 1896, nom. illeg. Types: China. Shaanxi, Thae-pei-san, Giraldi 996, 991, 992 (syntypes F).
Plants medium-sized to rather large, green or yellowish green, glossy, in densely or loosely caespitose mats. Stems prostrate or ascending, to 5–10 cm long, loosely subpinnately or irregularly branched, branchlets arcuate; stem oval in cross section; cortical cells small, quadrate or hexagonal; epidermal cells small, thick-walled, in 3–4 layers, distinctly hyaline; central strand weakly differentiated; pseudoparaphyllia large, numerous, foliose. Stem and branch leaves similar, falcate-secund, narrowly elongate-lanceolate, 3.5–5.0 mm × ca. 0.6 mm, plicate, gradually narrowed to a long acumen, involute to somewhat tubular; margins serrulate at upper part of leaf; costae slender, reaching 1/2 the leaf length or above, 40–80 mm wide at base; median leaf cells elongate-rhomboidal to linear, ca. 8–15 times as long as wide; basal cells broad, rectangular, thick-walled, porose; alar cells few, usually inflated, hyaline, thin-walled. Autoicous. Inner perichaetial leaves elongate-lanceolate, ca. 6.0 mm × 0.8–1.0 mm, plicate; costae single, slender. Setae 20–30 mm long, reddish brown; capsules erect, cylindrical, arching curved, variable in sizes to 1.5–3.0 mm long; annuli differentiated, in 2–3 rows; endostome segments carinate, hyaline, smooth at base, papillose above; cilia 1–2(–3), short, or as long as the segments, often nodulose. Spores 12–18 mm in diameter, papillose.


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