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Published In: Acta Botanica Yunnanica 3: 394. f. 4: 1–9. 1981. (Acta Bot. Yunnan.) Name publication detail

Project Name Data (Last Modified On 3/16/2009)
Acceptance : Accepted
Project Data     (Last Modified On 5/4/2009)
General Reference:

Illustration & Map     (Last Modified On 5/4/2009)
Illustrations: Grimmia mammosa C. Gao & T. Cao (Figs. 12–23.)
Map: Grimmia mammosa C. Gao & T. Cao (Distribution in China)

Distribution     (Last Modified On 5/4/2009)
Habitat: on exposed rocks or thin soil over rocks in high mountains; alt. 2500–3000 m.
Distribution: China, Bhutan, and Malawi (Muñoz 2000b).

Specimens Examined     (Last Modified On 5/4/2009)
Specimens Examined:
Chinese specimens examined: Xizang: Lang Co., M. Zang 1723 (HKAS); Ya-dong Co., S.-K. Chen 4983 (HKAS, IFSBH); A-sang District, M. Zang 116a (HKAS). Yunnan: Li-jiang Co., W.-X. Xu 0042 (ALTA, HKAS).

Notes     (Last Modified On 5/4/2009)
general taxon notes:
Grimmia mammosa is distinguished from other species of Grimmia by having bulging cells in the upper and middle part of the leaves, each with a distinct tuberculum. The costa at leaf base is very broad, up to 1/3 the leaf width and flattened in cross section, consisting of large, adaxial cells and an abaxial stereid band. The plants are rigid and dull and the leaves are unkeeled.
We have verified two specimens from Bhutan [D. G. Long 8787, 8296 (both in E)]. Even though only a few immature sporophytes were present in these specimens, they help our understanding of this species. Grimmia mammosa has erect, elongate setae, up to 4 mm long. The capsules are exserted, erect, ovoid to cylindrical and narrowed at the mouth. The sexual condition of the species is dioicous, and it appears to be distributed in the high mountains in the Himalayan regions. It is adapted to drought and high radiation environments and has extreme xerophytic features such as large bulging cells with intensely thickened walls and long hyaline awns.


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14. Grimmia mammosa C. Gao & T. Cao in C. Gao, G.-C. Zhang & T. Cao   粗瘤紫萼藓 cu-liu zi-e xian
Acta Bot. Yunnan. 3(4): 394. 1981. Type. China: Xizang (Tibet), Ya-dong Co., alt. 2900 m, M. Zang 71 (holotype IFSBH; isotypes ALTA, HKAS, NY).
Plants small, up to 1.5 cm high, dull, dark green to blackish green, in dense tufts. Stems sparsely forked, rigid, without distinct central strand in cross section. Leaves erect-appressed when dry, spreading when moist, 1.6–2.0 mm long, upper leaves larger than lower ones, broadly lanceolate to ovate-lanceolate, ending in a denticulate, hyaline hair-point (0.2–0.5 mm long), broadly concave, not keeled; the lower leaves blunt, muticous; margins plane, crenulate owing to bulging cells above middle part; costa broad, up to 1/3 the width of leaf base, becoming narrower toward apex, ending just below the awn, flattened, with a row of large adaxial cells and small abaxial stereid band in cross section; upper and median leaf cells bistratose, obscure, 5–7 µm wide, rounded-quadrate, distinctly bulging, each with a large tuberculum up to 4–5 µm in diameter; basal cells quadrate to subquadrate, 9–14 µm × 7–12 µm, yellowish, with thicker transverse walls. Dioicous. Perichaetial leaves similar to upper stem leaves, but with longer hair-points. Setae long, erect, up to 4 mm long; capsules exserted, erect, ovoid to cylindrical, narrowed at the mouth. Mature sporophytes not seen.
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