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Published In: Acta Botanica Fennica 131: 68. 1985. (Acta Bot. Fenn.) Name publication detail

Project Name Data (Last Modified On 3/6/2009)
Acceptance : Accepted
Project Data     (Last Modified On 3/6/2009)
General Reference:

Illustration & Map     (Last Modified On 3/6/2009)
Related Objects:

Click here for distribution map.

Illustrations: C. Gao (ed.) 1994 (Pl. 58, figs. 1–11).
Map: Campylopodium medium (Duby) Giese & J.-P. Frahm (Distribution in China)

Distribution     (Last Modified On 3/6/2009)


Habitat: on soil by road or in open fields.
Distribution: China, Japan, Indonesia, Malaysia, the Philippines, New Guinea, Pacific Islands, Australia, New Zealand, Chile, and Puerto Rico.

Specimens Examined     (Last Modified On 3/6/2009)
Specimens Examined:


Chinese specimens examined: Taiwan: Tai-pei Co., Chuang & Schofield 199, 204 (UBC), 203 (MO). Xizang: Mo-tuo Co., Y.-G. Su 1130 (KUN). Yunnan: Gong-shan Co., M.-Z. Wang 9693a (PE).


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1. Campylopodium medium (Duby) Giese & J.-P. Frahm  扭柄藓 niu-bing xian
Acta Bot. Fenn. 131: 68. 1985. Didymodon medius Duby in Moritzi, Syst. Verz. Zollinger Pfl. 134. 1846. Type. Indonesia: Java, Zollinger 411Za.
Aongstroemia euphoroclada C. Müll., Syn. Musc. Frond. 1: 429. 1848.
Campylopus euphorocladus (C. Müll.) Bosch & Sande Lac., Bryol. Jav. 1: 79. 1858.
Campylopodium euphorocladum (C. Müll.) Besch., Ann. Sci. Nat. Bot. sér. 5, 18: 189. 1873.
Dicranella euphoroclada (C. Müll.) Jaeg., Ber. Thätigk. St. Gallischen Naturwiss. Ges. 1870–71: 372. 1872.
Plants small, slender, 5–25 mm high, yellowish green to yellowish brown, sometimes shiny, in loose tufts. Stems short, erect, simple or sometimes branched, comose foliate in upper part; central strand slightly differentiated. Leaves 2–4 mm long, broadly ovate and sheathing at base, abruptly narrowed to a long subula; margins entire; costa occupying 1/4 of the leaf base width, filling all of the subula; upper shoulder cells short-rectangular to rhomboidal; lower shoulder cells narrowly rectangular; alar cells not differentiated. Dioicous. Perichaetial leaves not much differentiated. Setae up to 8 mm long, flexuose when dry, curved when moist; capsules ovoid, ca. 2 mm × 0.6 mm, with a short neck, sometimes slightly strumose, stomata superficial; annuli developed; opercula convex on the abaxial side, flat on the adaxial side 3 as long as capsules, obliquely rostrate; peristome teeth brownish-orange, striate below, hyaline or papillose above, divided halfway down. Calyptrae small, cucullate, entire at base. Spores 20–22 µm in diameter, smooth to finely papillose, yellowish brown.




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