Small to large shrubs and trees, unarmed, terrestrial, without raphides in the tissues, sometimes resinous at stem apices. Leaves opposite or rarely ternate, subsessile to petiolate, with tertiary and quaternary venation not lineolate, on the lower surface often with pubescent and/or foveolate domatia in the axils of the secondary veins; stipules quickly deciduous or rarely persisting, interpetiolar, shortly fused around the stem, or occasionally calyptrate (i.e, fused into a sheathing cap), triangular to ligulate or obovate, generally held erect and flatly pressed together in bud. Inflorescence terminal and in axils of the uppermost leaves, thyrsiform to paniculiform, multiflowered, pedunculate, with bracts developed to reduced. Flowers sessile to pedicellate, distylous, protandrous, medium to large, fragrant, apparently at least sometimes nocturnal; hypanthium ellipsoid to turbinate; calyx limb truncate to 5--7-lobed, without calycophylls; corolla salverform, white, glabrous or papillose internally in throat and on lobes, lobes 5--7, triangular, valvate in bud; stamens 5--7, inserted in corolla tube, with anthers narrowly ellipsoid, dorsifixed near base, included to partially exserted; ovary 2-locular, with ovules numerous in each locule, imbricated and ascending on axile placentas; stigmas 2-lobed, included or exserted. Fruit capsular, cylindrical to ellipsoid or ovoid, septicidally dehiscent from the base and/or sometimes from the apex, woody or infrequently chartaceous; seeds flattened, small, irregularly elliptic to oblong, marginally winged and often erose.