Shrubs and small to medium-sized trees, unarmed, terrestrial, without raphides in tissues, markedly resinous at stem apices, branching at least sometimes sylleptic. Leaves opposite, with tertiary and quaternary venation not lineolate but with numerous well developed intersecondary veins, on lower surface with well developed pubescence domatia at junctions of secondary veins with costa; stipules quickly deciduous, interpetiolar, triangular, generally held erect and flatly pressed together in bud. Inflorescences axillary (i.e., present in both axils of the node), borne with leaves near to well below stem apex, thyrsiform to paniculiform, multiflowered, pedunculate, bracteate. Flowers sessile or subsessile generally in groups of 3-5, small to medium-sized, distylous; hypanthium ellipsoid; calyx limb subtruncate to 5-lobed, without calycophylls; corolla funnelform to salverform, white, densely villous to villosulous internally in tube, in throat, sometimes on lobes, and on margins of lobes, lobes 5, triangular, valvate in bud; stamens 5, inserted in corolla tube, anthers narrowly ellipsoid, dorsifixed near base, included in long-styled flowers and exserted in short-styled flowers; ovary 2-locular, ovules numerous in each locule, imbricated and ascending on axile placentas; stigmas 2-lobed, rather succulent, exserted in long-styled flowers and included in short-styled flowers. Fruit capsular, cylindrical, woody, septicidally dehiscent from apex, valves spreading but remaining fused and persistent at base, sometimes a little asymmetric with capsule apex and entire calyx limb retained on one valve; seeds flattened, small, fusiform, marginally winged, entire to erose, acuminate at the ends or sometimes 2-lobed at one end.