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Published In: Chloris Melvilliana 9–10, pl. A. 1823. (Chlor. Melvill.) Name publication detail
 

Project Name Data (Last Modified On 7/20/2009)
 

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EUTREMA R. Brown, Chlor. Melvill., 193. 1823.

Ihsan A. Al-Shehbaz

Tribe: Eutremeae, Al-Shehbaz, Beilstein & E.A. Kellogg, Pl. Syst. Evol. 259. 112. 2006.

Name derivation: from Greek eu, well, and trema, opening, in reference to the perforation in the fruit septum.

Type species: E. edwardsii R. Brown.

Esquiroliella H. Léveillé, Mondes Plantes, ser. 2, 18: 31. 1916. Type species: E. violifolia (H. Léveillé) H. Léveillé (based on Martinella violifolia H. Léveillé).

Glaribraya H. Hara, J. Jap. Bot. 53: 135. 1978. Type species: G. lowndesii H. Hara.

Martinella H. Léveillé Bull. Soc. Bot. France 60: 290. 1904; not Martinella Baillon, Hist. Pl. 10: 30. 1891; not Martinella (Cooke & Massee ex Cooke) Saccardo, Syll. Fung. 10: 409. 1892. Type species: M. violifolia H. Léveillé.

Neomartinella Pilger in Engler & Prantl, Nat. Pflanzenfam. Nachtr. 3: 134. 1906. Type species: N. violifolia (H. Léviellé) Pilger (based on Martinella violifolia H. Léveillé).

Platycraspedum O. E. Schulz, Repert. Sp. Nov. Regni Veg. Beih. 12: 386. 1922. Type species: P. tibeticum O. E. Schulz.

Taphrospermum C. A. Meyer in Ledebour, Fl. Altaic. 3: 172. 1831. Type species: T. altaicum C. A. Meyer.

Thellungiella O. E. Schulz in Engler, Pflanzenreich IV. 105 (Heft 86): 251. 1924. Lectotype species designated by German (2002: ): T. salsuginea (Pallas) O. E. Schulz (based on Sisymbrium salsugineum Pallas).

Wasabia Matsumura, Bot. Mag. (Tokyo) 13: 71. 1899. Type species: Matsumura (1899) did not designate a type among his two species: W. pungens Matsumura and W. hederifolia (Franchet & Savatier) Matsumura. The former is illegitimate because it lists the earlier published Lunaria japonica Miguel as a synonym.

     Herbs, annual, biennial, or perennial, with slender or fleshy and fusiform roots, rhizomes, or caudex. Trichomes absent or simple. Multicellular glands absent. Stems erect or ascending to decumbent or prostrate, simple or branched at base and/or apically, leafy or rarely leafless. Basal

leaves petiolate, rosulate or not, simple, entire, dentate, or pinnately or palmately lobed; cauline leaves petiolate or sessile and cuneate or auriculate to amplexicaul at base, entire, dentate, or crenate, lowermost alternate or rarely verticillate; ultimate veins ending or not with apiculate callosities. Racemes bracteate throughout or basally, or ebracteate, elongated considerably or not in fruit; fruiting pedicels erect and subappressed to stem, ascending, or divaricate, persistent. Sepals ovate or oblong, free, deciduous or rarely persistent, erect, equal, base of inner pair not saccate; petals white or rarely pink to purple, longer or shorter than sepals, rarely absent; blade spatulate, obovate, oblong, or obcordate, apex obtuse, rounded to emarginate; claw undifferentiated from blade; stamens 6, exserted or included, tetradynamous to subequal in length, erect to spreading; filaments wingless or rarely flattened and laterally toothed, unappendages, glabrous, free; anthers ovate or oblong, obtuse or apiculate at apex; nectar glands lateral or confluent and subtending bases of all stamens, median glands present or absent; ovules 2–96 per ovary; placentation parietal. Fruit dehiscent, siliques or silicles, linear, oblong, ovoid, obcordate, conical, ovate, or lanceolate, terete, slightly 4-angled, latiseptate or angustiseptate, not inflated, unsegmented; valves papery, with an obscure or prominent midvein, glabrous or rarely papillate, smooth or torulose, not keeled, wingless; gynophore obsolete or to 5 mm; replum terete or flattened throughout or only basally; septum complete or rarely perforated and reduced to rim or absent, veinless; style obsolete or distinct and up to 3 mm, slender or clavate; stigma capitate, entire or rarely 2-lobed, unappendaged. Seeds uniseriate or biseriate, wingless, oblong to ovate, plump or flattened; seed coat obscurely reticulate to foveolate or papillate, slightly mucilaginous or not when wetted; cotyledons incumbent, oblique or rarely accumbent.

Twenty-six species: Eurasia, North America.

References: Al-Shehbaz (2000b, 2000i, 2007b), Al-Shehbaz & Warwick (2005), Al-Shehbaz et al. (2000b), Warwick et al. (2006a).

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