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Published In: Species Plantarum. Editio quarta 4(2): 637, 757[as Fluggea]–758. 1806. (Apr 1806) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: A. RADCLIFFE-SMITH
General/Distribution: A widespread genus of 13 species in the tropics and subtropics of both hemispheres, extending in places into the warm temperate zone. Represented in Pakistan by 2 species.


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Dioecious or rarely monoecious, unarmed or spinescent trees or shrubs, devoid of indumentum. Leaves alternate, shortly petiolate, stipulate, simple, entire, penninerved. Flowers axillary, usually in the proximal axils of the leafy branches, the males numerous, fasciculate, the females few or occasionally solitary. Male flowers: pedicels capillary; sepals 5, unequal, imbricate; petals 0; disc glands 5, interstaminal, alternating with the sepals; stamens 5, filaments free, anthers extrorse, longitudinally dehiscent; pistillode large, deeply 2-3-partite, the lobes often appendiculate or themselves bilobate. Female flowers: pedicels and sepals as in the ♂ flowers; petals 0; disc flat, annular, entire or shallowly 5-6-lobed; ovary smooth, 2-3-celled, with 2 ovules per cell; styles 2-3; connate at the base, recurved or spreading, bifid or bilobate. Fruit globose, slightly or scarcely fleshy, loculicidally dehiscent or rupturing irregularly; endocarp crustaceous, 3-celled or rarely 2 or 1-celled by abortion. Seeds 2 per cell, segmentiform, trigonous, smooth, ecarunculate; testa. thick, crustaceous; albumen scanty; embryo incurved; cotyledons ovate, elliptic or obovate, flat.

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1 A rigid shrub; branches pale grey or whitish, thorny; leaves rarely exceeding 2.5 cm, often obovate-spathulate, sometimes retuse or obcordate 2 Flueggea leucopyrus
+ A graceful shrub or small tree; branches usually reddish-brown, unarmed; leaves commonly more than 2.5 cm, obovate, rounded, obtuse or subacute 1 Flueggea virosa
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