Perennial herb, generally branched from base. Leaves opposite. Flowers in few flowered lateral cymes, rarely solitary. Calyx 5-partite, glandular within. Corolla subcampanulate, tube short, lobes 5, twisted to the right in bud. Corona epipetalous, arising a little above the origin of staminal column, lobes 5, generally broad, entire or bind, alternating with corolla-lobes. Staminal column short, anthers tipped by an inflexed membrane. Pollinia solitary in each cell, pendulous. Stigma exserted, fleshy, conical, clavate or peltate. Follicles usually echinate. Seeds comose.