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Published In: Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 66: 92. 1933. (Bot. Jahrb. Syst.) Name publication detail

Project Name Data (Last Modified On 7/20/2009)

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CUPHONOTUS O. E. Schulz, Bot. Jahrb. Syst. 66: 92. 1933.

Ihsan A. Al-Shehbaz


Name derivation: from Greek cuphos, Light weight, and notus, back, in reference to the poorly developed keel or the valve back.

Lectotype species (designated by Shaw (1974: 154): C. humistratus (F. Meuller) O. E. Schulz

     Herbs annual. Trichomes absent or simple, usually crisped. Multicelluar glands absent. Stems erect to decumbent, branched. Basal leaves subpetiolate, subfleshy, not forming a rosette, simple, entire or shallowly pinnatifid; cauline leaves subsessile and cuneate to attenuate, not auriculate at base, entire, dentate, or pinnatifid. Racemes many-flowered, ebaracteate, corymbose, elongated considerably in fruit; rachis straight; fruiting pedicels divaricate, persistent. Sepals oblong to elliptic, free, deciduous, spreading, equal, base of inner pair not saccate; petals white or creamy white, spreading, longer to subequaling sepals; blade oblanceolate to linear, apex obtuse, margin obscurely lobed; claw obscurely distinct from blade, shorter than sepals, glabrous, unappendaged, entire; stamens 6, exserted, spreading, tetradynamous; filaments wingless, unappendaged, glabrous, free, dilated at base; anthers ovate, rounded at base, obtuse at apex; nectar glands 4, distinct; median nectaries absent; lateral nectaries 1 on each side of lateral stamens; ovules 2-15 per ovary; placentation parietal. Fruit dehiscent, capsular silicles, ovoid, ellipsoid, to subglobose, slightly angustiseptate, not inflated, sessile, unsegmented; valves subleathery, veinless or prominently veined, glabrous, not keeled and rounded, smooth, wingless, unappendaged; gynophore absent; replum rounded; septum complete, membranous, without a midvein; style obsolete or short and less than 1 mm long, stout, persistent, glabrous; stigma capitate, entire or slightly 2-lobed, unappendaged. Seeds uniseriate, margined, ellipsoid, sligtly flattened; seed coat smooth, copiously mucilaginous when wetted; cotyledons incumbent, entire.

Two species; Australia.

Reference: Shaw (1974).



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1. Plants pubescent with simple trichomes; fruit valves smooth; petals 1-1.5 mm ........  1. C. andreanus

1. Plants glabrous; fruit valves conspicuously reticulate; petals 2-3 mm ....................  2. C. humistratus


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