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  • Reference article Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera. 2012. Verbenaceae. 4(2): 453–473. In G. Davidse, M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera (eds.) Fl. Mesoamer.. Missouri Botanical Garden Press, St. Louis.
  • Reference article Lorence, D. H. 1999. A nomenclator of Mexican and Central American Rubiaceae. Monogr. Syst. Bot. Missouri Bot. Gard. 73: 1–177. View in BotanicusView in Biodiversity Heritage Library
  • Reference article Standley, P. C. & L. O. Williams. 1975. Rubiaceae. 24(11): 1–274. In P. C. Standley & L. O. Williams (eds.) Fl. Guatemala, Part XI, Fieldiana, Bot.. Field Museum of Natural History, Chicago.
 
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