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Published In: Species Plantarum 2: 1054. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: A. RADCLIFFE-SMITH
General/Distribution: A pantropical but predominantly Neotropical genus of 110 species, with only 1 species in S. Pakistan.


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Monoecious shrubs, subshrubs or woody herbs, mostly twining. Indumentum simple and sometimes urticating. Leaves alternate, petiolate, stipulate, stipellate, simple, 3-5-lobate or - foliolate, entire of toothed. Inflorescences bisexual, axillary or terminal, usually solitary and long-pedunculate, crowded and subtended by 2 large stipulate outer bracts; outer bracts equal or subequal, entire, toothed or trilobate, green or coloured; male partial inflorescence a pedunculate pleiochasium of 5(-7) sessile (1-) 3-flowered cymes together with a mass of fused aborted flowers, all surrounded by 4 free or fused inner bracts; female partial inflorescence a subsessile 2-3-bracteate (1-) 3-flowered dichasium abaxial to the male structure. Male flowers: pedicels articulate; buds globose, later splitting into 4-6 valvate lobes; petals 0; disc 0; stamens 7-90, commonly 15-30, filaments usually fused into a column, free portions short, anthers longitudinally dehiscent. pistillode 0. Female flowers: sepals 5-12, imbricate, narrow, rarely entire, usually pinnatifid, sometimes glandular, in fruit accrescent, indurated and enclosing the fruit; petals 0; disc usually 0; ovary 3(-4)-celled, with 1 ovule per cell; styles completely fused, elongate, obtuse, dilated or obliquely excavated at the apex. Fruit trilobate, dehiscing into 3 bivalved cocci. Seeds globose, ecarunculate; albumen fleshy; cotyledons broad, flat.
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