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Published In: Species Plantarum 1: 18. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
General/Distribution: About 250 species, chiefly American; represented here by 4 species.


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Herbs with opposite, decussate, rarely whorled leaves. Inflorescence spicate, rarely pseudumbellate or fascicled, terminal, rarely axillary. Flowers bisexual, small or moderate in size, bracteate. Calyx tubular, 5-ribbed, unequally shortly 5-toothed, not enlarging in fruit. Corolla salver-shaped, 5-lobed, somewhat bilabiate and lobes subequal, oblong to obovate. Stamens usually 4, didynamous, not exserted; anthers ovate, connective with or without a glandular appendage. Carpels 2, syncarpous; ovary entire to 4-lobed, 4-loculed with 1 ovule in each locule; style reaching the level of stamens. Fruit schizocarpic, enclose ed within the calyx, separating into four, 1-seeded nutlets; seeds oblong, angular.

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1 Spikes lax and elongated, especially in fruit (2)
+ Spikes densely flowered, paniculately orymbose (3)
2 (1) Leaves deeply serrate, lyrate pinnatifid or subentire. Corolla-tube c. 6 mm long Verbena officinalis
+ Leaves bipinnatisect with narrowly linear segments. Corolla-tube. c. 10 mm long Verbena tenuisecta
3 (1) Leaves oblong-lanceolate with amplexicaul bases, sharply serrate. Flowers c. 5 mm across. Calyx-tube 3-4 mm long Verbena bonariensis
+ Leaves oblong-ovate or ovate-triangular, without amplexicaul bases, pinnatifidly incised or coarsely serrate. Flowers c. 8 mm across. Calyx-tube c. 10 mm long Verbena incisa
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