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Published In: Bijdragen tot de flora van Nederlandsch Indië 12: 603. 1825. (Bijdr. Fl. Ned. Ind.) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: A. RADCLIFFE-SMITH
General/Distribution: A genus of 7 species extending from Pakistan to China and Indonesia (Sumbawa). One species only in Pakistan.


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Monoecious or dioecious erect, few-branched shrubs, subshrubs or woody herbs with a simple indumentum. Leaves alternate, petiolate, stipulate, simple, sinuate-dentate or lobate, 3-5-nerved from the base, otherwise penninerved, with a pair of glands at the base of the lamina. Inflorescences axillary, unisexual or bisexual, racemose or paniculate, dense or lax, ♂ many-flowered, ♀ few-flowered. Male flowers: sepals (4-) 5(-6), ± orbicular, imbricate; petals 0; disc annular, lobate or composed of (4-) 5(-6) free glands; stamens 10-30, on a convex receptacle, the outer opposite the sepals, filaments free, anthers introrse, longitudinally dehiscent, connective broad; pistillode minute or 0. Female flowers: sepals 5-6, imbricate, sometimes toothed, sometimes accrescent ;petals 0; disc annular, entire; ovary 3-celled, with 1 ovule per cell; styles 3, stout, recurved, shortly bifid. Fruit trilobate, dehiscing septicidally and loculicidally into 6 valves; endocarp woody. Seeds ovoid, testa crustaceous, albumen fleshy; cotyledons broad, flat.
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