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Published In: Species Plantarum 1: 218. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
General/Distribution: 47 species; distributed in Europe, Mediterranean region to montane regions of W. Pakistan, India and North and South Africa.


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Annual or perennial, much branched, prostrate or decumbent, hairy or subglabrous rarely glabrous, mat forming herbs. Leaves opposite, the upper ones often alternate through abortion of one member of a pair, small, sessile or subsessile, entire, obovate to oblanceolate or narrowly elliptic, hairy or sub-glabrous, rarely entirely glabrous; stipules interpetiolar, small, scarious, ovate-triangular with ciliate margins. Inflorescence dense leaf opposed or pseudoaxillary rarely terminal cymose sessile fascicles or clusters. Flowers sessile or subsessile, tetra or pentamerous, perigynous, hairy throughout or subglabrous to glabrous at perigynous zone, hairs straight or uncinate; bracteate, bracts inconspicuous. Sepals 4-5, plano-concave, with or without distinct fimbriate membranous margins, unequal or subequal to equal in length, connivent to erect. Petals 4-5, inconspi¬cuous, subulate-filiform, inserted on rim of cupule, alternate with and shorter than the sepals. Stamens 2-5, antisepalous, inserted on rim of cupule, filaments short, filiform, anthers squarish-oblong or ovoid. Carpels 2, syncarpous; ovary ovoid to sub- globose, glabrous, smooth or papillose in the upper part, membranous, unilocular, ovule 1, basal, amphitropous; style very short, bifid, rarely 2 or absent. Fruit shorter than to slightly exceeding the sepals, 1-celled, 1-seeded, ovoid¬subglobose or ovate-ellipsoid, smooth or papillose in the upper part, membranous, indehiscent utricle enclosed by the persistent calyx. Seed shining dark brown or black, crustaceous, lenticular to ovate-obvoate, smooth; embryo annular, enclosing farinaceous endosperm.

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1 Sepals 5, equal or slightly unequal, margins distinctly membranous. Stamens 3-5. Perigynous zone with straight hairs or almost glabrous (2)
+ Sepals 5, unequal (2 outer long, 3 inner short), margins not membranous. Stamens 2, opposite to 2 outer long sepals. Perigynous zone always with uncinate hairs Herniaria cinerea
2 (1) Annual herb. Flowers 1-1.5 mm long, sessile. Sepals. mostly equal or somewhat subequal in older flowers. Perigynous zone subglabrous to almost glabrous Herniaria hirsuta
+ Perennial herb. Flowers 1.8-2.5 mm long, sub-sessile. Sepals always equal. Sepals and perigynous zone covered with straight hairs Herniaria cachemiriana
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