Small to medium-sized shrubs, unarmed, terrestrial, with raphides in tissues, generally with many or all of the leaves borne on short-shoots (brachyblasts), generally with rather large pustlose to pellucid glands in the tissues. Leaves opposite, often small, with tertiary and quaternary venation not lineolate, without domatia, sometimes (Heterophyllaea pustulata) with margins crenate or serrate; stipules interpetiolar or shortly fused around stem or to petiole bases, generally persistent with leaves, triangular, generally erect and flat or perhaps imbricated in bud. Inflorescences terminal and in uppermost leaf axils, in few-flowered lax cymes or with flowers solitary, pedunculate, bracteate. Flowers distiylous, rather large and showy, pedunculate or pedicellate, sweetly fragrant, perhaps sometimes nocturnal; hypanthium ellipsoid; calyx limb deeply 5-lobed, without calycophylls; corolla salverform, white to dark blue, purple, or violet, pubescent internally in tube with pubescence pattern apparently differing between flower morphs, externally generally glabrous, lobes 5, triangular, valvate in bud; stamens 5, inserted in upper part of corolla tube, filaments short, anthers narrowly ellipsoid, dorsifixed, included in long-styled flowers and partly to fully exserted in short-styled flowers; ovary 2-locular, ovules numerous in each locule, imbricated on axile placentas; stigmas 2-lobed, exserted in long-styled flowers and included in short-styled flowers. Fruit capsular, subglobose to weakly didymous, stiffly chartaceous to woody, loculicidally dehiscent from apex, valves remaining fused at base for a time, spreading up to ca. 45°, with the suborbicular septum and attached placentas generally separating and falling from capsule, with calyx limb persistent; seeds flattened, medium-sized, suborbicular to ellipsoid, marginally winged, entire to finely densely erose.