DILOPHIA Thomson, Hooker’s J. Bot. Kew Gard. Misc. 5: 19. 1853.
Ihsan A. Al-Shehbaz
Name derivation: from Greek dis, double, and lophia, appandage or crest, in reference to the two crests at the fruit apex.
Type species: Dilophia salsa Thomson.
Herbs, perennial; roots fleshy, conical-linear, apically scaly. Trichomes absent or simple. Multicellular glands absent. Stems erect, simple underground, often branched aboveground. Basal leaves sessile, fleshy, rosulate, simple, entire or sinuate to dentate; cauline leaves sessile, attenuate, not auriculate at base, entire. Racemes bracteate or ebracteate, several- or many-flowered, dense, in simple or compound corymbs, not elongated in fruit; rachis straight; fruiting pedicels slender, terete, ascending to divaricate. Sepals broadly ovate, free, persistent, ascending, glabrous or puberulent, equal, base of inner pair not saccate. Petals white to lavender, erect at base with flaring blade, longer than sepals; blade obovate to spatulate, apex obtuse to emarginate, margin entire; claw obscurely to distinctly differentiated from blade, broad and sometimes as wide as blade, glabrous, unappendaged, entire. Stamens 6, tips slightly exserted, somewhat spreading, slightly tetradynamous; filaments filiform, wingless, unappendaged, not dilated at base, glabrous, free; anthers ovate, subobtuse at base, distinctly apiculate at apex, not coiled after dehiscence. Nectar glands 1, confluent and subtending bases of all stamens; median nectaries present; lateral nectaries semiannular. Ovules 4-12 per ovary; placentation parietal or subapical. Fruit dehiscent, capsular silicles, obcordate, strongly flattened and angustiseptate, not inflated, sessile, unsegmented; valves membranous or thin papery, obscurely veined, glabrous or puberulent, not keeled and rounded, smooth, erect or divergent on the broadly expanded base of replum, wingless, apically gibbous and cristate; gynophore absent; replum strongly flattened, base much broader, rim at ca. 45° angle to horizontal septum complete or perforated, reduced and to ca. 1 mm long, tanslusent, veinless; style to 0.5 mm long, cylindric, persistent, glabrous, included in apical notch of fruit; stigma capitate, entire, unappendaged. Seeds aseriate, wingless, broadly oblong to ovoid, slightly flattened; seed coat minutely reticulate, not mucilaginous when wetted; cotyledons inccumbent or oblique, entire.
Two species: Bhutan, China, India, Kyrgyzstan, Nepal, Pakistan, Tajikistan.