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  • Reference article Davidse, G., M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera. 2012. Rubiaceae a Verbenaceae. 4(2): i–xvi, 1–533. In G. Davidse, M. Sousa Sánchez, S. Knapp & F. Chiang Cabrera (eds.) Fl. Mesoamer.. Missouri Botanical Garden Press, St. Louis.
  • Reference article Forzza, R. C. 2010. Lista de espécies Flora do Brasil http://floradobrasil.jbrj.gov.br/2010. Jardim Botânico do Rio de Janeiro, Rio de Janeiro.
  • Reference article Idárraga-Piedrahita, A., R. D. C. Ortiz, R. Callejas Posada & M. Merello. (eds.) 2011. Fl. Antioquia: Cat. 2: 9–939. Universidad de Antioquia, Medellín.
  • Reference article Lorence, D. H. 1999. A nomenclator of Mexican and Central American Rubiaceae. Monogr. Syst. Bot. Missouri Bot. Gard. 73: 1–177. View in BotanicusView in Biodiversity Heritage Library
  • Reference article Molina Rosito, A. 1975. Enumeración de las plantas de Honduras. Ceiba 19(1): 1–118.
  • Reference article Standley, P. C. & L. O. Williams. 1975. Rubiaceae. 24(11): 1–274. In P. C. Standley & L. O. Williams (eds.) Fl. Guatemala, Part XI, Fieldiana, Bot.. Field Museum of Natural History, Chicago.
  • Reference article Stevens, W. D., C. Ulloa Ulloa, A. Pool & O. M. Montiel. 2001. Flora de Nicaragua. Monogr. Syst. Bot. Missouri Bot. Gard. 85: i–xlii,.
  • Reference article Taylor, C.M. 2010-2020. Studies in Neotropical Rubiaceae.
 
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