Climbing herbs or small shrubs, glabrous or pubescent. Leaves mostly cordate, entire, dentate or lobed. Flowers axillary, solitary, in scorpioid cymes or umbelliform or head-l ike cymes. Sepals equal or unequal, mostly pubescent. Corolla campanulate or funnel-shaped, blue, lilac or white, deeply lobed, dentate or entire. Style 1, stigmas 2, ellipsoid or oblong and complanate. Fruit capsular, globose to subglobose, with 4 major valves, these sometimes splitting further into 8. Seeds glabrous, tuberculate, winged or pilose.