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Published In: Species Plantarum 2: 866. 1753. (1 May 1753) (Sp. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library
 

Project Name Data (Last Modified On 1/2/2012)
Acceptance : Accepted
Project Data     (Last Modified On 1/2/2012)
Contributor Text: M. Qaiser and Rubina Abid
Contributor Institution: Federal Urdu University of Arts, Science & Technology, Karachi.

and

Department of Botany, University of Karachi, Karachi-75270

General/Distribution: A large genus with c. 1250 species of worldwide distribution (except Antarctica); represented in Pakistan by 15 species.

 

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Annual or perennial herbs, undershrubs, shrubs or small trees, generally with alternate leaves, quite variable in habit and foliage. Capitula radiate or discoid, rarely solitary axillary, usually few to numerous in terminal simple or compound corymbs or thryses. Involucre calyculate. Florets mostly yellow, sometimes pink, purple or violet, rarely white. Ray florets, usually conspicuous, sometimes minute. Disc florets 3-many. Anthers ecaudate, shortly auriculate, filaments collar dilated, slightly to markedly dilated towards the base, endothecial tissues radial. Style branches truncate to obtuse with a crown of hairs without appendages with discrete stigmatic lines. Cypselas ellipsoid or ovoid or oblong ribbed, glabrous or pubescent, pappus bristles many, uniform.
 

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1.
+
Capitula radiate or sometimes few, inconspicuous ray florets.
 
 
3
 
 
Capitula discoid without marginal radiate female florets.
 
 
2
 
2.
+
Capitula in dense corymbs. Phyllaries 18-22; outer bracteoles 7-11 with long blackish tips.
 
 
 
9. S. vulgaris
 
 
Capitula in lax corymbs. Phyllaries c. 15, outer bracteoles 4-5, without or sometimes with short blackish tips.
 
 
 
11. S. dubitabilis
 
3.
+
Plants perennial.
 
4
 
 
Plants annual.
 
11
 
4.
+
Ray florets minute, inconspicuous, shorter than the style.
 
 
1. S. graciliflorus
 
 
Ray florets conspicuous, at least 4 mm long, longer than the style.
 
 
5
 
5.
+
Cypselas 6-8 mm long, pappus bristles 10-14 mm long.
 
 
10. S. paulsenii
 
 
Cypselas (1.5-) 2-4 (-4.5) mm long, pappus bristles (3.5-) 4-7 (-8) mm long.
 
 
6
 
6.
+
Basal and cauline leaves entire to crenate-dentate.
 
 
7
 
 
Basal and at least some of the cauline leaves pinatipartite to pinnatisect or lyrate.
 
 
 
8
 
7.
+
Stem leafy, basal leaves long petiolate. Cauline leaves many. Phyllaries 15-21.
 
 
12. S. pseudo-orientalis
 
 
Stem subscapose, basal leaves sessile to subsessile. Cauline leaves few. Phyllaries 12-13.
 
 
 
14. S. nudicaulis
 
8.
+
Phyllaries 8. Ray florets 4-5, ligule 3-4 mm long.
 
 
2. S. royleanus
 
 
Phyllaries 10-21. Ray florets 6-20, ligule 6.5-12 mm long.
 
 
9
 
9.
+
Phyllaries 16-21, lanceolate, acumi-nate. Ray florets 16-20. Capitula 2-6.
 
 
3. S. tibeticus
 
 
Phyllaries 10-13, oblong-ovate or oblong lanceolate often with black tip. Ray florets 6-15. Capitula 6-15 to numerous.
 
 
 
10
 
10.
+
Involucre 3-5 x 6-8 mm. Capitula in dense corymbs. Ray florets 7-13 mm long.
 
 
 
13. S. analogus
 
 
Involucre 5-8 x 10-18. Capitula in lax corymbs. Ray florets 12-18 mm long.
 
 
4. S. korshinskyi
 
11.
+
Ray florets short, scarcely exceeding the involucre.
 
 
12
 
 
Ray florets conspicuous, 6 mm long, much exceeding the involucre.
 
 
5. S. glaucus
 
12.
+
Leaves entire to denticulate, not lobed or pinnatipartite.
 
 
6. S. flavus
 
 
Leaves pinnatipartite, pinnatifid or irregularly lobed.
 
 
13
 
13.
+
Plants glabrous, tall herbs, up to 60 cm high.
 
 
15. S. ramosus
 
 
Plants subglabrous to glabrecent, dwarf herbs, up to 20 cm tall.
 
 
14
 
14.
+
Capitula narrowly cylindrical, peduncle up to 1 cm long.
 
 
7. S. krascheninnikovii
     
 
Capitula campanulate, peduncle 1.5-4 cm long.
 
 
8. S. ladakhensis
 

 
 
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