YINSHANIA Y. C. Ma & Y. Z. Zhao, Acta Phytotax. Sin. 17(3): 113. 1979.
Ihsan A. Al-Shehbaz
Name derivation: Yinshan Mountain in Neimongol and Gansu provinces (China).
Type species: Yinshania albiflora Y. C. Ma & Y. Z. Zhao (=Y. acutangula (O. E. Schulz) Y. H. Zhang).
Cochleariella Y. H. Zhang & R. Vogt, Acta Bot. Bor.-Occid. Sin. 9(4): 224. 1989. Type species: C. zhejiangensis (Y. H. Zhang) Y. H. Zhang & R. Vogt (based on Cochleariopsis zhejiangensis Y. H. Zhang.
Cochleariopsis Y. H. Zhang, Acta Bot. Yunnan. 7(2): 143. 1985; not Löve & Löve, Bot. Notiser 128: 513. 1976. Type species: C. zhejiangensis Y. H. Zhang & R. Vogt.
Hilliella (O. E. Schulz) Y. H. Zhang & H. W. Li, Acta Bot. Yunnan. 8: 401. 1986; Cochlearia L. sect. Hilliella O. E. Schulz, Notizbl. Bot. Gart. Berlin-Dahlem, 8: 544. 1923. Type species: H. fumarioides (Dunn) Y. H. Zhang & H. W. Li (based on Cochlearia fumarioides Dunn).
Herbs annual or perennial with elongated or rarely tuberous rhizomes. Trichomes absent or present and simple or 1- or 2-forked. Multicellular glands absent. Stems erect to ascending, sometimes decumbent, simple or branched basally and/or apically. Basal leaves petiolate, not forming a rosette, simple, undivided to pinnately or palmately lobed, trifoliolate, or pinnately, compound; cauline leaves petiolate, not auriculate, similar to basal leaves but progressively smaller and with less divisions or simple upward. Racemes few- to many-flowered, ebracteate or bracteate, corymbose, dense or lax, elongated in fruit; rachis straight, or rarely strongly flexuous; fruiting pedicels erect and subappressed to rachis, ascending, divaricate, or reflexed, persistent. Sepals oblong or rarely ovate, free, deciduous, spreading, equal, base of inner pair not saccate; petals white or rarely pinkish, spreading, longer than sepals; blade obovate or rarely oblong or ovate, apex rounded; claw not or obscurely differentiated from blade, glabrous, unappendaged, entire; stamens 6, spreading, subequal in length; filaments filiform, wingless, unappendaged, not dilated at base, glabrous, free; anthers ovate or oblong, rounded at base, not apiculate at apex; nectar glands 4, 1 on each side of lateral stament, median nectaries absent; ovules 1-24 per ovary; placentation parietal. Fruit dehiscent, capsular silicles, oblong, ovoid, ellipsoid, or globose, terete or slightly flattened and latiseptate, rarely slightly angustiseptate, not inflated, sessile, unsegmented; valves papery, veinless or obscurely veined, glabrous, papillose, or with simple hairs, not keeled, smooth, wingless, unappendaged; gynophore absent or ca. 0.5 mm; replum rounded; septum complete, perforated, or absent, membranous, not veined; style obsolete to 3 mm, persistent; stigma discoid, entire, unappendaged. Seeds 1-12 per locule, aseriate, uniseriate, or biseriate, wingless, oblong, ovate, reniform, orbicular, ovoid, lenticular, elliptic, ellipsoid, or globose, sligtly flattened; seed coat minutely reticulate or papillate, not mucilaginous when wetted; cotyledons incumbent or rarely accumbent, entire. n = 6, 7, 21.
Thirteen species: endemic to China, one species also each in Taiwan and Vietnam.
References: Al-Shehbaz et al. (1998), Koch & Al-Shehbaz (2000), Zhang (2003).
see interactive key to the species