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Published In: Exposition des Familles Naturelles 2: 123. 1805. (Feb-Apr 1805) (Expos. Fam. Nat.) Name publication detail

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: Ghulam Rasool Sarwar
Contributor Institution: Department of Botany, University of Karachi, Karachi.
General/Distribution: A family of c. 33 genera and c. 1500 species distributed all over the world except Occeania. Represented in Pakistan by 9 genera and 32 species.


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Annual, biennial or perennial herbs, rarely undershrubs or shrubs, usually succulent, sometimes woody at base. Leaves alternate, opposite or verticillate, usually simple, sometimes compound, exstipulate, often thick and fleshy, sometimes forming a basal rosette. Flowers actinomorphic, (3-)4-5(-32)-merous, usually bisexual, in axillary or terminal cymes, often grouped in corymb- or panicle-like inflorescences, less often in racemes or spike or solitary in the leaf axil. Sepals free or united. Petals free or united, sometimes forming a tubular corolla. Stamens as many or twice as many as the petals, filaments free or more or less adnate to the corolla; anthers 2-celled, usually basifixed, introrse, dehiscing longitudinally. Carpels superior or semi-inferior, equal in number to the petals, free or united at the base. Nectar scales usually present, small, applied to the base of the outer face of the carpels. Fruit a follicle. Seeds few-numerous, very small with smooth, rugose or tuberculate tegument and usually fleshy endosperm and straight embryo.

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1 Stamens as many as petals. (2)
+ Stamens twice as many as petals. (3)
2 (1) Leaves opposite, connate at the base. Flowers minute in the axils of the leaves 0.9-2.5 mm long, 3-5 merous. Tillaea
+ Leaves alternate, not connate at the base. Flowers small, terminal 3-6 mm long, 3-9-merous. Sedum
3 (1) Leaves decussate. Flowers 4-merous, large, petals forming tubular corolla. (4)
+ Leaves not decussate. Flowers 3-9-merous, small, petals free or connate basally till the middle. (5)
4 (3) Sepals fused, calyx tubular and inflated. Stamens inserted at the base of corolla. Flowers pendent. Bryophyllum
+ Sepals free or basally united. Stamens inserted at or above middle of corolla. Flowers erect on pedicel. Kalanchoe
5 (3) Plants with scaly leaves. (6)
+ Plants without scaly leaves. (7)
6 (5) Petals free or nearly free. Rhodiola
+ Petals connate nearly to the middle. Pseudosedum
7 (5) Ovaries stipitate – attenuate at the base. (8)
+ Ovaries sessile, not attenuate at the base. (9)
8 (7) Inflorescence a raceme. Petals basally connate. Basal cauline leaves forming rosettes. Orostachys
+ Inflorescence corymbose. Petals free. Basal cauline leaves not forming rosettes. Hylotelephium
9 (7) Plants with basal rosette. Petals united. Style situated on the dorsal side of ovary. Rosularia
+ Plants without basal rosette. Petals free or basally united. Styles situated on the ventral side of ovary. (10)
10 (9) Foliage radical leaves forming rosette. Rhizome thick. Rhodiola
+ Foliage radical leaves absent. Rhizome absent or thin. Sedum
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