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Published In: Genera Plantarum 76. 1789. (4 Aug 1789) (Gen. Pl.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: Y. NASIR
Contributor Institution: Stewart Herbarium, Gordon College, Rawalpindi.
General/Distribution: A family of 50 genera and about 500 species, mostly temperate and tropical; especially in S. Africa, Australia, Mediterranean region; less common in S. America and the Pacific Islands. Represented in W. Pakistan by 5 genera and 7 species.
Comment/Acknowledgements: Acknowledgements: We are grateful to the United States Department of Agriculture for financing this research under P.L. 480. To Dr. R.K. Brum¬mitt (Kew) and Mr. B.L. Burtt (Edinburgh) for their help and to Dr. B. Peterson (Goteborg) for going through the manuscript and giving valuable suggestions and supplying a list of plants from foreign herbaria for citation. We are also indebted to the Herbarium Director, Pakistan Forest Research Institute, Peshawar for the loan of plant specimens.


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Perennials, a few herbs or shrublets. Leaves entire, alternate or opposite, usually numerous, exstipulate. Flowers rarely solitary, in racemes, spikes or fascicles; bisexual, regular. Calyx tubular, 4-5 lobed; lobes imbricate. Petals small or absent. Stamens as many as or double the number of lobes, episepalous. Carpels 1-2, rarely more, syncarpous; ovary unilocular, with one laterally attached pendulous ovule; style simple. Fruit a berry or a drupe. Seeds solitary.

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1 Calyx tube without disc scales (2)
+ Calyx tube with disc scales (4)
2 (1) Shrubs or herbs up to 60 cm tall. Fruit dry (3)
+ Shrubs up to 2.5 m tall. Fruit succulent Daphne
3 (2) Annual herbs. Flowers ebracteate, in lax terminal racemes. Calyx tube constricted in fruit Diarthron
+ Perennials, rarely annual. Flowers bracteate, axillary, solitary or in clusters. Calyx tube not constricted in fruit Thymelaea
4 (1) Disc unilateral, linear to lanceolate, some times 2-fid. Flowers in terminal clusters. Calyx tube splitting transversely above the ovary Stellera
+ Disc of 2-4 linear scales, rarely less. Flowers in terminal short racemes or spikes. Calyx tube not splitting above the ovary Wikstroemia
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