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Published In: Flora Aegyptiaco-Arabica 38. 1775. (Fl. Aegypt.-Arab.) Name publication detailView in BotanicusView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Contributor Text: YASIN J. NASIR
Flower/Fruit: Fl. Per.: December-April.
Type: Type: Arabia: Felix Lohaja, Forsskal (C).
Distribution: Distribution: N. Africa, Arabia, Pakistan.
Comment/Acknowledgements: In the present work a broad view has been taken of A. bacciferum. The species is very variable in habit, leaf and fruit characters and in the past, several infraspecific taxa have been recognised (Kazmi, l.c. 161); The characters employed to separate the taxa are too relative and variable. For instance Kazmi (l.c.) distinguishes ssp. bacciferum from ssp. lignosum, in the nutlets being either broad or thin margined and their degree of separation at maturity; further Kazmi's recognition of 4 varieties is not satisfactory. The var. tuberculosum, although generally with narrower leaves, may in some cases exhibit leaves up to 10 mm broad (A. Rahman 25721; Sultanul Abedin 4251). From specimens seen of (Heliotropium tuberculosum), it appears that this taxon frequents coastal areas and is characterised by its short twisted stems and branches and small leaves. Insect galls are frequent on the branches. It is doubtful if the corky or membranous back of the nutlets (Jafri, 1966, p. 281) is of any taxonomic value. Hooker (l.c.) on the basis of this character described a new variety: Heliotropium undulatum var. suberosa. This corky nature occasion-ally occurs in M. crispum and to a varying degree in Heliotropium bacciferum (M. Salim 43). In the form of leaves and habit, as already pointed out by Kazmi (l.c. p. 164) the var. lignosum resembles var. tuberculosum and var. fartakense bears similarity; with var. bacciferum. In my view Heliotropium bacciferum has been too finely split into infraspecific forms. As already noted, intermediate forms occur which cannot with certaintly be ascribed to any particular taxon discussed above. The species is allied to Heliotropium crispum.
Illustration: Heliotropium bacciferum Forssk. (Illustration)
Map Location: Sind: Karachi: G-4 Karachi, T. Cooke 8 (K); id., H. Rich 437 (K); id., M. Nath & G. D. Samson 15320 (RAW); Manora, M. Salim 43 (RAW); Cliffden, A. Rahman 25721 (RAW); id., 5.4.1964 M. A. Siddiqi (RAW); Sonmiani, Y. Nasir 8619 (RAW); Cape Monze-Paradise Point, Sultanul Abedin 4219 (KUH, RAW); Karachi-Bela, 34th mile, on sandy hill, Sultanul Abedin 2154 (KUH, RAW); Mangopir, S. I. Ali 334 (KUH); id., M. Salim 57 (RAW); Karachi University Campus, 1962 Q. Sultana s.n. (KUH); Clifton, Jafri 1396; 1401 (E); id., coll. ignot. (PPFI- B); Maripur, along Lyari, Jafri 889 (E); near Bihar Colony, Jafri 889 (KUH); P. E. C. H. S., 10.1.1957, R. Islam s.n. (KUH); Karachi beach, 19.5.1955, Kitamura s.n. (KUH); Gharo, 2.5.1964, S. I. Ali s.n. (KUH); Intelligence School, 20.11.1959, S. I. Ali s.n. (KUH); National Refinery, S. I. Ali & Sultanul Abedin 660 (KUH); Sonmiani, S. A. Farooqi & Sultanul Abedin 1208 (KUH); near Bihar Colony, Jafri 889 (KUH); Damloti-Khadeji, Rech. f. 28511 (M); Dabeji, 35 miles E. of Karachi, Burtt 1566 (E); Baluchistan: beyond Hub River, R. R. Stewart 29112 (RAW); G-2 Kappar-Gwadur, Rech. f. 27898 (B, K, M, US); Sunstar-Gwadur, Ghafoor 6327 (KUH); Pasni-Kappar, Rech. f. 27867 (M); Pasni-Turbat, 17th mile, A. Ghafoor & M. Qaiser 173 (KUH); Uthal-Naka, Kharai, Rech. f. 27474 (M, K); 30-50 km W. of Bela, Rech. f. 27600 (M); 10-15 km S. of Sehwan, Rech. f. 28739 (M); Ras Nuh to Gwadur, J. Lamond 483 (E); Gwadur-Pasni, Sultanul Abedin & A Hussain 6433 (KUH); Pasni, S. I Ali et at 739; 741 (KUH); Sunster Rest House, S. I. Ali et at 1127 (HUH); Pasni-Kappar, J. Lamond 465 (E); 3 miles from Gwadur, S. I. Ali et al 1145 (KUH); Locust R. House, Pasni, M. Qaiser et al 955 (KUH).


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Perennial, decumbent or procumbent with a woody base, short and stout up to 15 mm thick at base. Leaves lanceolate to linear-lanceolate, 5-55 x 2-16 mm, flat or t terete, hairy on both surfaces, margin revolute, longer hairs stiff, up to 2 mm long, arising from bulbous base. Inflorescence reduced, up to 20 mm long, bearing close set uniseriate flowers. Calyx persistent, 2-2.5 mm long, 5-partite into lanceolate lobes, hairy to the outside; hairs often stiff and arising from a bulbous base. Corolla white, c. 2.5-3 mm long; tube shortly cylindrical, hairy outside, glabrous within; lobes imbricate, 0.7-0.8 mm long, oblong to suborbicular, crenulate to t undulate. Anthers 1-1.2 mm long, elongate, broader at base, attached c. 1.1 mm from the corolla base, slightly furrowed and sometimes 2-fid at apex. Style shorter than stigma. Fruit globose, usually hairy when young; nutlets 4, brown, margin winged, back rugulose, sometimes with a membranous inflated back.
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