Lateral nectar glands semiannular or annular, open towards the inner side, rarely in pairs or short; middle glands often joining the laterals, torose and ± thickened in the middle, rarely short, conical and not joining the laterals. Hairs simple or branched, sometimes absent, often rigid and ciliate, rarely glandular. Sepals erect or suberect, Stamen filaments not appendaged but sometimes broad at base; anthers usually oblong, obtuse. Stigma depressed capitate, often sub-bilobed. Fruits usually long (siliquae) rarely short (siliculae) ; valves often somewhat convex and rigid; style mostly short, rarely absent; stigma mostly broad, obscurely to sub-bilobed; septum usually complete, rarely perforated or absent,with or without a vein, rarely 2-4-veined; seeds usually oblong or suborbicular, rarely winged; mucilaginous or not when wet; radicle mostly incumbent.