Tropicos | Name - Boechera grahamii (Lehm.) Windham & Al-Shehbaz

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Boechera grahamii (Lehm.) Windham & Al-Shehbaz

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40. Boechera grahamii (Lehmann) Windham & Al-Shehbaz, Harvard Pap. Bot. 12: 241. 2007; Turritis grahamii Lehmann, Del. Sem. Hort. Hamb. 7. 1831 (also in Linnaea 6: Litt. 74. 1831); based on T. patula Graham, Edinburgh New Philos. J. [7]: 350. 1829, not T. patula Ehrhart, Beitr. 7: 159. 1792; Arabis patula Torrey, Bot. U.S. and Mex. Bound. Surv. 1: 32. 1859, not A. patulaWeinmann, Cat. Hort. Dorp. 18. 1810, nor A. patula Host, Fl. Austr. 2: 271. 1831. TYPE: Rocky Mts., Drummond s.n. (Lectotype designated by Windham & Al-Shehbaz (2007: 241), K!).

Turritis brachycarpa Torrey & A. Gray, Fl. N. Amer. 1: 79. 1838; Arabis drummondii A. Gray var. brachycarpa (Torrey & A. Gray) A. Gray, Manual, ed. 5. 69. 1867; A. confinis S. Watson var. brachycarpa (Torrey & A. Gray) S. Watson & J. M. Coulter in A. Gray, Manual, ed. 6. 67. 1890; A. brachycarpa (Torrey & A. Gray) Britton, Mem. Torrey Bot. Club. 5: 174. 1894, not Ruprecht, Fl. Caucas. 73. 1869; A. holboellii Hornemann var. brachycarpa (Torrey & A. Gray) S. L. Welsh, Utah Fl., ed. 3. 256. 2003; Boechera brachycarpa (Torrey & A. Gray) Dorn, Vasc. Pl. Wyoming, ed. 3. 375. 2001; Turritis drummondii (A. Gray) Lunell var. brachycarpa (Torrey & A. Gray) Lunell, Amer. Midl. Naturalist 5: 236. 1919; TYPE: United States, Michigan, Gratiot Co., Fort Gratiot [Point Huron], Z. Pitcher s.n. (lectotype designated by Holmgren (2004: 245), NY!).

Arabis confinis S. Watson, Proc. Amer. Acad. Arts 22: 466. 1887. TYPE: Canada, Manitoba, Lake Winnipeg Valley, 1857, E. Bourgeau s.n. (lectotype designated by Windham & Al-Shehbaz (2007: 241), GH!; isolectotype, E!).

Arabis bourgovii Rydberg, Mem. N.Y. Bot. Gard. 1: 186. 1900. TYPE: Same as Turritis patula Graham.

Arabis dacotica Greene, Leafl. Bot. Observ. Crit. 2: 80. 1910; A. divaricarpa A. Nelson var. dacotica (Greene) B. Boivin, Amer. Midl. Nat. 54: 510. 1955. TYPE: United States, South Dakota, Fort Meade, W. H. Forwood 28 (holotype, US!).

Arabis divaricarpa A. Nelson var. stenocarpa M. Hopkins, Rhodora 39: 133. 1937; A. patula Torrey var. stenocarpa (M. Hopkins) Farwell, Papers Mich. Acad. Sci. 26: 14. 1941. TYPE: Canada, Quebec, Rimouski Co., slaty ridges east of the village, Bic, 4 Jul 1907, M. L. Fernald & J. F. Collins 1057 (holotype, GH!; isotype, CAN!).

Arabis divaricarpa var. hemicylindrica B. Boivin, Amer. Midl. Nat. 54: 510. 1955; A. boivinii G. A. Mulligan, Rhodora 97: 157. 1996. TYPE: Canada, Saskatchewan, District de Maple Creek, 10 miles au sud, Carmichael, monts Cypres, ecorre de la coulee du ruisseau Bone, 9 Jul 1952, B. Boivin & J. F. Alex 9738 (holotype, DAO!; isotypes, GH!, MT).

Plants short-lived perennials to biennials, with or without caudices, lacking crowded, persistent leaf bases; apomictic, with spheroid pollen. Stems usually 1 per caudex branch, arising near ground surface from center of basal rosettes, (1.2–)2.0–12.0 dm, lower parts glabrous or with simple and sessile or subsessile, 2–3(–6)-rayed trichomes to 1 mm, upper parts glabrous. Leaves at stem bases oblanceolate, 1.5–10(–20) mm wide, entire to dentate, ciliate proximally with simple trichomes to 1 mm, blade surfaces sparsely to densely pubescent with sessile or subsessile, 2–4(–7)-rayed trichomes 0.2–0.6 mm; cauline leaves (10–)13–52, often concealing stem proximally, the uppermost glabrous, with auricles 1–5 mm. Inflorescences mostly unbranched, 16–88-flowered; fruiting pedicels 5–22(–30) mm, divaricate-ascending to descending, gently curved downward or rarely straight, glabrous. Flowers divaricate at anthesis; sepals glabrous or pubescent; petals white or rarely lavender, 5.5–8.0 ´ 1.0–2.0 mm, glabrous; ovules 84–146 per fruit. Fruits 3.5–9.0 cm ´ 1.0–1.8 mm, divaricate-ascending to pendent, not appressed to rachises, not secund, straight or slightly curved, with parallel edges, glabrous; style 0.05–0.5 mm. Seeds uniseriate to subbiseriate, 1.0–1.8 ´ 0.8–1.2 mm; wing continuous, 0.1–0.25 mm wide. 2n = 21.

Flowering: May–Aug.

Habitat: rocky slopes and sandy soil in prairies and open forests.

Elevation: 100–3150 m.

Distribution: Canada (Alberta, British Columbia, Manitoba, Northwest Territories, Ontario, Quebec, Saskatchewan, Yukon), United States (Alaska, Colorado, Michigan, Minnesota, Montana, New York, Dakota, South Dakota, Vermont, Washington, Wisconsin, Wyoming).

Reproductive mode: apomictic triploid.