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Published In: The Flora of British India 6(20): 731–732. 1894. (Fl. Brit. India) Name publication detailView in Biodiversity Heritage Library

Project Name Data (Last Modified On 6/2/2011)
Acceptance : Accepted
Project Data     (Last Modified On 6/3/2011)
Synonym Text: C. nigerrima Nelmes in Kew Bull.: 200. 1939; C. atrata L. subsp. pullata (Boott) Kük. in Engler, Pflanzenr. IV.20, 38: 400. 1909; C. atrata L. var. pullata Boott, l.c. 3: 114. 1862.
Flower/Fruit: Fl. Per.: July–August.
Type: Syntypes: India, Gurwhal, 13-14000 ft., Duthie 4499; Sikkim, 11-15000 ft, J.D.Hooker s.n. (K).
Habitat: High mountain meadows, 2800-5000 m.
Distribution: Distribution: From N Pakistan to Sikkim and Central China.
Comment/Acknowledgements: Description above is based primarily on the type specimen of C. nigerrima. C. duthiei is also closely related to C. perfusca V. Krecz., differing in having acute, papillose female glumes as long as or longer than utricles, without scarious margins. The constancy of these characters could not be verified as I have seen very little material. However, it should be noted that C. perfusca has been variously treated in relation to C. atrata L. (Kukkonen in Rech.f., l.c. 277. 1998).
Map Location: B-7 Swat: Bishigram Lake, 10-11000’, Stewart & Rahman Beg 24968 (K); Hazara: Kagan Valley, Saif ul Muluk, Ali s.n. (RAW); B-8 Kashmir: Tragbol Pass, 12000 ft, Stewart 4920 (RAW); Burzil Pass, Clarke 29722 (K); Sind Valley, Ward in Duthie 17825 (K); Khillanmarg, Stewart 8595a (K).


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Caespitose perennial, 40-60 cm. Rhizome short, horizontal. Stem sharply trigonous, smooth or slightly scabrous above, papillose. Leaves shorter than stem; sheaths 40-100 mm, reddish brown, strongly nerved, glossy, scarious side brown-spotted, margin concave; ligule to 0.7 mm, white with brownish spots, arch higher than wide; blades 2.5-3.5 mm wide, flat, keeled, margins revolute, both sides papillose, margins and keel scabrous. Inflorescence of 3-4(-5) spikes. Bracts equalling or overtopping inflorescence, with sheath to 5 mm. Terminal spikes to 24 mm, club-shaped, gynecandrous; other spikes 15-22 x 5-6 mm, ellipsoid or cylindrical, female or gynecandrous, peduncle to 45 mm; female glumes 3.9-4.3 x 1.5-2 mm, ovate, acute, dark reddish brown, papillose; utricles 4-4.2 x c. 1.9 mm, elongate ellipsoid, yellowish or golden-brown, with reddish brown spots, glossy, papillose, beak 0.1-0.2 mm, cylindrical, truncate, dark reddish brown. Stigmas 3. Nut c. 2.3 x 1.3 mm, including 0.4 mm persistent style base and a 0.1 mm stipe, widely obovoid, trigonous, light brown, reticulate with papilla in each areole.
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