Macromitriaceae Churchill in Churchill & Linares, Bibliot. José Jerónimo Triana 12: 588. 1995.
Plants small to robust, in cushions, tufts, or mats on rocks or trees. Stems erect, erect‑ascending or creeping with erect‑ascending branches, densely or sparsely tomentose. Leaves crowded, often hygroscopic, erect to crispate or spirally twisted when dry, spreading to squarrose when wet, more or less keeled, lanceolate, ovate‑lanceolate, oblong‑lanceolate, ligulate, or lingulate; margins plane or revolute, rarely incurved, generally entire, but often serrate or denticulate; costa single, strong, ending near the apex; upper leaf cells generally small, irregularly rounded and thick‑walled (less frequently elongate), papillose or smooth, basal leaf cells generally elongate less frequently short and rounded, tuberculate or smooth, alar cells undifferentiated, weakly differentiated, rarely strongly differentiated. Brood bodies sometimes present in leaf axils, on rhizoids or leaves. Dioicous, autoicous, or synoicous. Perichaetia usually terminal, occasionally lateral. Setae smooth or roughened. Capsules immersed, emergent, or exserted, erect and symmetric, smooth or furrowed; opercula rostellate or rostrate; stomata present, immersed or superficial; peristome diplolepidous, teeth none, single or double, preperistome sometimes present, exostome teeth 8 or 16, endostomial basal membrane present or absent, endostome segments absent, 8 or 16, narrow, non‑perforate, alternating with teeth (rarely appearing opposite), cilia none. Spores isosporous or anisosporous, sometimes multicellular. Calyptrae campanulate, conic, or cucullate, naked or hairy, plicate or non‑plicate.