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Published In: Kryptogamenflora der Mark Brandenburg, Laubmoose 675, 692. 1905. (Krypt.-Fl. Brandenburg, Laubm.) Name publication detail

Project Name Data (Last Modified On 3/30/2009)
Acceptance : Accepted
Project Data     (Last Modified On 3/30/2009)
General Reference:

Notes     (Last Modified On 3/30/2009)
general taxon notes:
Three species of Helodium were reported in China (C. Gao et al. 1977). Two species were accepted by Crosby et al. (2000) because Helodium sachaliense was placed in the genus Bryonoguchia. Although Helodium blandowii (Web. & Mohr) Warnst. was reported from northeastern China (C. Gao et al. 1977), we could not confirm the report. Two species, H. paludosum (Austin) Broth. and H. sachalinense (Lindb.) Broth. were treated by P.-C. Wu et al. (2002) in the Thuidiaceae. However, H. paludosum (Austin) Broth. was treated in the Helodiaceae (He 2008) and H. sachalinense (Lindb.) Broth. was treated under Echinophyllum of the Thuidiaceae. Echinophyllum sachalinense is clearly distinct from the remaining species of Helodium (O’Brien & Horton 2000).



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8. Helodium Warnst. 沼羽藓属 zhao-yu xian shu
Krypt.-Fl. Brandenburg, Laubm. 2: 675, 692. 1905, nom. cons.
Thuidium sect. Elodium (Sull.) Braithw., Brit. Moss. Fl. 3: 8. 1896. Hypnum subg. Elodium (Sull.) Lesq. & Jam., Man. N. Amer. Moss. 329. 1884. Thuidium subg. Elodium (Sull.) Lindb., Musc. Scand. 31. 1879. Elodium (Sull.) Aust., Musci Appalach. 52. 1870. Hypnum sect. Elodium Sull. in Gray, Manual (ed. 2), 668. 1856.
Plants rather robust, green or yellowish green, rarely brownish, not shinning or slightly shinning, in tufts or cushions. Stems erect or ascending, simple or forkedly branched, but usually regularly pinnately branched; paraphyllia dense, filamentous, simple or branched, pale, brownish when old; central strand absent. Stem leaves and branch leaves similar, gradually narrowed from an ovate-lanceolate or cordate base to a lanceolate apex, longitudinally plicate, restricted at base; margins entire or serrate, revolute above, long ciliate or smooth at base; costa rather rigid, reaching upper portion of leaf apex; leaf cells hyaline, elongate-hexagonal or oval-hexagonal, basal cells rectangular. Autoicous or dioicous. Inner perichaetial leaves erect, lanceolate, deeply plicate, pale; costa ending below leaf apex. Setae slender, smooth; capsules ovoid or cylindrical, strongly recurved or horizontal, brownish; annuli present, stomata present at hypophysis; opercula conic; exostome teeth yellowish brown; cilia weak, usually 3. Calyptrae smooth. Spores not seen.

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1. Plants robust; stem leaves oblong at the base, gradually lanceolate toward the apex; leaf margins entire or papillose; leaf cells papillose at upper corner; costa smooth on the back.............................. 1. H. paludosum
1. Plants rather small; stem leaves cordate at the base, suddenly narrowed to a ciliate apex; basal leaf margin multiplicate; leaf cells centrally papillose; costa papillose or setulosely papillose on the dorsal surfaces....
...............................................................................................................................2. H. sachalinense
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