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Published In: Bryologia Universa 2: 610. 1827. (Bryol. Univ.) Name publication detail

Project Name Data (Last Modified On 4/2/2009)
Acceptance : Accepted
Project Data     (Last Modified On 4/2/2009)
General Reference:

Illustration & Map     (Last Modified On 4/2/2009)
Related Objects:
Illustrations: P.-C. Wu & Y. Jia (eds.) (2004), Pl. 64, figs. 1–10.

Click here for distribution map.

Map: Hypnum hamulosum Froel. (Distribution in China)

Distribution     (Last Modified On 4/2/2009)
Habitat: on soil, humus, rocks, grassland, or rarely on tree trunks; alt. 1200–4800 m.
Distribution: China, Russia, Europe, and North America.

Specimens Examined     (Last Modified On 4/2/2009)
Specimens Examined:
Chinese specimens examined: Anhui: Mt. Huang, D.-K. Li 2423, 2432 (both in SHM). Gansu: Tian-zhu Co., S.-Z. Hu 524 (PE). Guizhou: Jiang-kou Co., Mt. Fanjing, Guiyang Teachers’ Univ. Biol. Dept. 45, 40069a, 50323 (all in PE); Sui-yang Co., C. Gao & J.-Y. Feng 003307 (IFSBH). Hebei: Huo-shi valley, Q.-W. Wang 1177 (PE). Heilongjiang: Da Xing An Ling (Greater Khingan Mt.), C. Gao 2714, 12712, 12982, 13099, 13196, 17095 (all in IFSBH); Wu-ying Natural Reserve, P.-C. Chen & C. Gao 598 (IFSBH); Xiao Xing An Ling (Lesser Khingan Mt.), C. Gao 619 (IFSBH). Henan: Mt. Laojun, J.-X. Luo 235 (MO, PE); Xin-yang City, Li-jia-Cai, W.-M. Xiong 645 (PE). Hubei: Mt. Shennongjia, X.-W. Zhang s.n. (IFSBH). Hunan: Mt. Heng, G. Yang 8 (MO, PE). Jiangsu: Su-zhou City, Dongshan (Mt.), M. Zang s.n. (HKAS). Jiangxi: Mt. Lu, X.-X. Zhong & X.-Z. Sun 20a (WH). Jilin: Mt. Changbai, C. Gao 22275, 22325 (both in IFSBH), C. Gao 22120 (MO). Nei Mongol: Da Xing An Ling (Greater Khingan Mt.), Gen-he Co., D.-Z. Huang 352 (PE); Mt. Helan, Z.-G. Tong 2141 (PE); Mt. Wangfeng, P.-C. Chen & C. Gao 812, 987 (both in IFSBH). Shaanxi: Chang-an Co., M.-X. Zhang & J.-Z. Zhang 1948a (XBGH); Mt. Taibai, S.-E. Liu & B.-Q. Zhong 4335 (XBGH), X.-J. Li 652 (PE); Ning-shan Co., M.-X. Zhang & Y.-G. Wang 3944 (XBGH). Sichuan: A-ba Co., S. He 31040 (MO, PE); Ba-tang Co., S. He 31626 (MO, PE); Da-jin Co., 8th Forest Team 4823 (PE); Dao-cheng Co., X.-J. Li 81-3109 (HKAS); Lei-bo Co., Z.-T. Guan 8746 (PE); Li-xian Co., X.-J. Li 1877 (HKAS); Ma-bian Co., Q.-L. Zhang 101101 (PE); Mt. Omei, C. Gao 18804, 18825, 18828 (all in IFSBH); Ma-er-kang Co., S. He 31271a (MO); Mu-li Co., M.-Z. Wang 23558 (PE); Nan-ping Co., S. He 30358 (MO, PE); Ruo-er-gai Co., S. He 30765 (MO, PE); Song-pan Co., S. He 30423a (PE); Tian-quan Co., Mt. Erlong, J.-L. Yang 2010 (HKAS); Xiang-cheng Co., X.-J. Li 81-3067 (HKAS); Ya-jiang Co., S. He 31973 (MO, PE). Xinjiang: Manasi, X.-S. Wang 6 (PE); Mt. Altai, J.-C. Zhao s.n. (XBGH); Mt. Tianshan, X.-S. Wang 33 (PE); Mu-lei Co., S.-L. Chen 560b (MO). Xizang: Cha-yu Co., M.-Z. Wang 8267, 8440c (both in PE), L.-S. Wang 1153 (HKAS); Chang-du City, M. Zang 5289a (HKAS); Cuo-na Co., Y.-C. Yang 45 (PE); Gong-bu-jiangda Co., W.-H. Li 2154b (HKAS); Ji-long Co., S.-K. Chen 240, 249 (both in HKAS); Lin-zhi Co., K.-Y. Lang 302b (PE); Mi-lin Co., S.-K. Chen 182 (HKAS), Y.-G. Su 5292 (HKAS); Ya-dong Co., M. Zang 134, 208, 216, 457, 527 (all in HKAS); Zha-mu Co., S.-G. Wu 1127a (HKAS); Zou-gong Co., M.-Z. Wang 14648 (PE). Yunnan: Da-li Co., S.-E. Liu 21136 (HKAS); Gong-shan Co., M.-Z. Wang 11568a (PE), M. Zang 3117 (HKAS), D.-C. Zhang 2416 (HKAS); Jiang-cheng Co., L.-S. Wang 81-2583 (HKAS); Li-jiang Co., X.-J. Li 85-294 (HKAS), Mt. Jade Dragon, M. Zang 1647, 1648 (both in HKAS), W.-X. Xu 1000 (PE); Mt. Meilishi, M. Zang 84566 (HKAS); Wei-xi Co., L.-S. Wang 82-428 (HKAS), D.-C. Zhang 705 (HKAS); Zhong-dian Co., L.-S. Song 81-1187 (HKAS), X.-J. Li 81-1026 (HKAS), H.-F. Xie 1967 (MO, PE). Zhejiang: Mt. West Tian-mu, M. Zang 562 (HKAS).

Notes     (Last Modified On 4/2/2009)
general taxon notes:
Hypnum hamulosum is similar to H. recurvatum (Lindb. & Arn.) Kindb., but differs in having slenderly acuminate leaf apices, less developed alar cells, and inflated, hyaline stem epidermal cells, which are very thin-walled and easily broken when dry.


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9. Hypnum hamulosum Schimp. in B.S.G.   弯叶灰藓   wan ye hui xian
Bryol. Eur. 6: 96, tab. 590 (fasc. 57–61, Mon. 20). 1854. Drepanium hamulosum (Schimp.) Jens., Meddel. Gronland 3: 326. 1887. Type: Europe.
Hypnum cupressiforme var. hamulosum Brid., Muscol. Recent. Suppl. 2: 217. 1812.
Plants small, rather slender, soft, yellowish green, somewhat glossy, in dense mats. Stems prostrate, or erect-ascending when growing in dense mats, ca. 3 cm long; in stem cross section a layer of enlarged, hyaline, thick-walled, epidermal cells, central strand poorly developed; regularly or irregularly pinnately branched, branches flattened, 3–5 mm long; pseudoparaphyllia few, lanceolate, at base of branches. Stem leaves strongly falcate-secund, 1.4–1.8 mm × 0.5–0.7 mm, broadly elliptical-lanceolate or ovate-lanceolate, slenderly acuminate; margins plane, entire or serrulate above the midleaf, sometimes revolute below; costae short, double or absent; leaf cells linear, median cells 39–80 µm × 3–5 µm, basal cells oblong-elliptical, sometimes porose, slightly yellowish; alar cells few or indistinctly developed, quadrate or hexagonal, hyaline; branch leaves smaller, similar to stem leaves in shape. Dioicous or autoicous with separate perichaetia and perigonia. Inner perichaetial leaves plicate. Setae elongate, 1.2–2.0 cm long, yellowish or reddish, twisted when dry; capsules cylindrical, 1.5–2.5 mm long, suberect to horizontal, slightly arcuate, orange-yellow, smooth, constricted below the mouth when dry; opercula conic; annuli developed, in 2 rows of cells; peristome typically hypnoid, with 2 cilia. Calyptrae not seen. Spores 14–16 µm in diameter, brownish green, papillose.
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