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Published In: Commentario della Società Crittogamologica Italiana 2(3): 283. 1867. (Comment. Soc. Crittog. Ital.) Name publication detail

Project Name Data (Last Modified On 4/3/2009)
Acceptance : Accepted
Project Data     (Last Modified On 5/6/2009)
General Reference:

Illustration & Map     (Last Modified On 5/6/2009)
Illustrations: Ptilium crista-castrensis (Hedw.) De Not. (Figs. 1–9.)
Map: Ptilium crista-castrensis (Hedw.) De Not. (Distribution in China)

Distribution     (Last Modified On 5/6/2009)
Habitat: on humus, soil near bogs or streams, also on rocks, rotten wood, or tree trunks in forests or alpine bushes; alt. 1200–4600 m.
Distribution: China, Japan, Korea, Mongolia, Bhutan, Nepal, Sikkim, Russia, Europe, and North America.


Specimens Examined     (Last Modified On 5/6/2009)
Specimens Examined:
Chinese specimens examined: Guizhou: Jiang-kou Co., Mt. Fanjing, C. Gao & J.-Y. Feng s.n. (IFSBH). Hebei: Dong-ling, L.-Q. Yu 495 (PE). Heilongjiang: Da Xing An Ling (Greater Khingan Mt.), H.-F. Xie 95 (IFSBH), C. Gao 12708, 13197 (both in IFSBH); De-Du Co., C. Gao 13294 (MO); Wei-he Co., X.-W. Wang 143 (MO); Mu-ling Co., C. Gao 13581 (IFSBH, MO); Ning-an Co., C. Gao 6710 (IFSBH, MO); Xiao Xing An Ling (Lesser Khingan Mt.), Wu-ying, C. Gao 159, 3854 (both in IFSBH, MO). Hubei: Shennongjia, Q.-C. Li 10 (XBGH). Jiangxi: Yi-feng Co., Nanjing Univ. 72 (XBGH). Jilin: An-tu Co., S.-E. Liu 3838, 4002 (both in IFSBH, MO), C. Gao 33884 (IFSBH); Ji-an Co., C. Gao 1142 (MO); Mt. Changbai, S.-E. Liu  1520 (IFSBH, MO), C. Gao 1138, 22201 (both in IFSBH, MO), Vitt 34761 (ALTA, MO); Wu-song Co., S.-E. Liu 4313 (MO). Liaoning: Zhang-wu Co., S.-E. Liu 3104 (MO). Nei Mongol: Da Xing An Ling (Greater Khingan Mt.), Mt. Wufeng, C. Gao 3639 (IFSBH, MO), C. Gao 949, 1020 (both in IFSBH, MO); Tong-liao City, S.-E. Liu 3127 (IFSBH, MO). Shaanxi: Mt. Taibai, Z.-C. Zhu 7 (XBGH). Shanxi: Ning-wu Co., C. Gao 31001 (IFSBH). Sichuan: Ba-tang Co., S. He 31549a, 31563d (both in MO, PE); Dao-cheng Co., S. He 31728b, 31838 (both in MO, PE), M.-Z. Wang 81-4243a (PE); Dao-fu Co., Z.-T. Guan 60154 (MO); Ma-bian Co., F.-Z. Wang 2402 (PE); Ma-er-kang Co., S. He 31159 (MO, PE), X.-J. Li 1163 (HKAS); Mu-li Co., Y.-L. Zhang s.n. (IFSBH), P.-Q. Li 42151 (PE), M.-Z. Wang 23621b (PE), L.-S. Wang 83-1013, 83-1147, 83-1270 (all in HKAS); Song-pan Co., S. He 30485 (MO); Xiang-cheng Co., S. He 31749a (MO), X.-J. Li 3055 (HKAS), L.-S. Wang 81-2433 (HKAS); Ya-jiang Co., S. He 32011 (MO); Yan-yuan Co., M.-Z. Wang 2284 (PE), L.-S. Wang 83-956 (HKAS). Taiwan: Tai-chung Co., Shevock 18024, 18028 (both in CAS, MO). Xinjiang: Ha-mi Co., J.-C. Zhao 4536 (XJU); Urum-qi Co., S. He 951028, 951089 (both in MO). Xizang: Bo-mi Co., J.-W. Zhang 1502a (HKAS), Y.-G. Su 935 (HKAS); Cha-yu Co., J.-W. Zhang 7310, 7306a (both in HKAS); Ding-jie Co., K.-Y. Lang 1121b (PE); Dong-ga-la, M. Zang 332 (HKAS); Ji-long Co., W.-H. Li 75-369 (HKAS), S.-K. Chen 246 (HKAS); Lang-xian Co., M. Zang 1878, 1882, 1904, 1908 (all in HKAS); Lin-zhi Co., S.-K. Chen 339, 341 (both in HKAS), K.-Y. Lang 301 (PE); Lu-lang Station, W.-H. Li s.n. (HKAS); Mi-lin Co., S.-K. Chen 157, 196, 4889a (all in HKAS), K.-Y. Lang 390, 425, 581, 609, 613 (all in PE), M. Zang 1925, 1931, 1959 (all in HKAS), Y.-G. Su 5227, 5273, 5285, 5318 (all in HKAS); Mo-tuo Co., S.-G. Wu 5169, 5174 (both in HKAS); Mt. Songta, M.-Z. Wang 8276, 12282b, 12779b (all in PE); Mt. Sula, M. Zang 8082, 8108, 8121 (all in HKAS); Ri-dong Co., M. Zang 5098, 5599b, 5615b (all in HKAS); She-ji-la, S.-K. Chen 366, 411 (both in HKAS); Ya-dong Co., M. Zang 156, 215, 228, 398, 581 (all in HKAS); Zou-gong Co., D.-C. Zhang 7501 (HKAS). Yunnan: Bin-chuan Co., M.-K. Cui s.n. (YUKU); Da-li Co., Redfearn & Y.-G. Su 1350 (MO); De-qin Co., L.-S. Wang 81-2382 (HKAS); Fu-gong Co., M.-Z. Wang 6745 (PE); Gong-shan Co., M.-Z. Wang 1136b, 7808, 8072, 8218 (all in PE), A.-L. Zhang 51 (HKAS); Kunming City, S.-D. Wu & Z.-F. Fang 32 (PE); Li-jiang Co., S.-G. Wu 184 (HKAS); Wei-xi Co., M.-Z. Wang 5053, 5059a (both in PE); Yang-bi Co., Redfearn et al. 849 (MO); Zhong-dian Co., X.-J. Li 81-828, 81-845, 81-918, 81-1069 (all in HKAS), H.-F. Xie 1986 (PE).


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1. Ptilium crista-castrensis (Hedw.) De Not.   毛梳藓   mao shu xian
Cronac. Briol. Ital. 2: 17. 1867. Ctenium cristacastrensis (Hedw.) Jens., Danmarks Mosser 2: 122. 1923. Hypnum crista-castrensis Hedw., Sp. Musc. Frond. 287. 1801. Type: Europe.
Plants pale green or yellowish green, somewhat glossy, in dense mats. Stems prostrate, 5–15 cm long, densely pinnately branched, branches flattened; pseudoparaphyllia narrowly lanceolate. Stem leaves and branch leaves dimorphic; stem leaves broadly ovate-triangular, 2–3 mm × 0.8–1.2 mm, gradually acuminate, strongly squarrose or curled; margins serrulate above; costae double, reaching up to midleaf; leaf cells linear; alar cells few, only consisting of several quadrate or shortly rectangular cells; branch leaves narrowly ovate-lanceolate, falcate-secund, costae not distinct. Dioicous. Setae reddish brown, 1.5–2.5 cm long; capsules inclined to horizontal, oblong-ovoid, 2.0–2.5 mm long, arcuate when dry; opercula conic, bluntly and shortly rostrate; annuli 1–2 rows of cells; peristome teeth normal. Calyptrae not seen. Spores not seen.


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